FAGACEAE(Flora of china)壳斗科

FAGACEAE(Flora of china)壳斗科
FAGACEAE(Flora of china)壳斗科

FAGACEAE

壳斗科 qiao dou ke

Huang Chengjiu (黄成就 Huang Ching-chieu)1, Zhang Yongtian (张永田 Chang Yong-tian)2; Bruce Bartholomew3 Trees or rarely shrubs, monoecions, evergreen or deciduous. Stipules usually early deciduous. Leaves alternate, sometimes false-whorled in Cyclobalanopsis. Inflorescences unisexual or androgynous with female cupules at the

base of an otherwise male inflorescence. Male inflorescences a pendulous head or erect or pendulous catkin, sometimes branched; flowers in dense cymules. Male flower: sepals 4–6(–9), scalelike, connate or distinct; petals absent; filaments filiform; anthers dorsifixed or versatile, opening by longitudinal slits; with or without a

rudimentary pistil. Female inflorescences of 1–7 or more flowers subtended individually or collectively by a cupule formed from numerous fused bracts, arranged individually or in small groups along an axis or at base of an androgynous inflorescence or on a separate axis. Female flower: perianth 1–7 or more; pistil 1; ovary inferior, 3–6(–

9)-loculed; style and carpels as many as locules; placentation axile; ovules 2 per locule.Fruit a nut. Seed usually solitary by abortion (but may be more than 1 in Castanea,Castanopsis,Fagus, and Formanodendron), without endosperm; embryo large.

Seven to 12 genera (depending on interpretation) and 900–1000 species: worldwide except for tropical and S Africa; seven genera and 294

species (163 endemic, at least three introduced) in China.

Many species are important timber trees. Nuts of Fagus,Castanea, and of most Castanopsis species are edible, and oil is extracted from nuts of Fagus. Nuts of most species of this family contain copious amounts of water soluble tannin. Members of the Fagaceae are the main element of

both broad-leaved evergreen and mixed mesophytic forests from 500–3200 m.

Huang Chengchiu, Chang Yongtian, Hsu Yongchun & Jen Hsienwei. 1998. Fagaceae. In: Chun Woonyong & Huang Chengchiu, eds. Fl. Reipubl. Popularis Sin. 22: 1–332.

1a. Male inflorescences heads, pendulous; female flowers (1 or)2; germination epigeal ................................ 1. Fagus

1b. Male inflorescences elongated catkins, erect or pendulous; female flowers solitary or in spikes;

germination hypogeal (epigeal in Formanodendron).

2a. Male inflorescences erect; anthers ca. 0.3 mm; stigma punctiform or a minute terminal pore.

3a. Leaves deciduous; ovary 6(–9)-loculed ..................................................................................... 2. Castanea 3b. Leaves evergreen; ovary 3(–6)-loculed.

4a. Cupules solitary on rachis, mostly bilaterally symmetric with abaxial and adaxial sides

differing in shape and/or spines (± radially symmetric in C. calathiformis,C. cerebrina,C.

uraiana,

and C. sclerophylla) ........................................................................................................ 3. Castanopsis 4b. Cupules in cymes on rachis (often some cupules abort but still evident at base of developed cupules, except in L. fenzelianus, radially symmetric ...................................................... 4. Lithocarpus 2b. Male inflorescences pendulous; anthers 0.5–1 mm; stigma capitate or dilated.

5a. Cupules splitting into 3–5 valves; nuts angular, winged .............................................. 5. Formanodendron 5b. Cupules indehiscent; nuts circular in cross section, wingless.

6a. Bracts of cupule not united, arranged spirally ......................................................................6. Quercus

6b. Bracts of cupule united, arranged in concentric rings .............................................. 7. Cyclobalanopsis

1. FAGUS Linnaeus, Sp. Pl. 2: 997. 1753.

水青冈属 shui qing gang shu

Trees deciduous. Bark smooth. Winter buds elongated, reddish brown, apex acute. Leaves alternate, 2-ranked,

plicate in bud; secondary veins ± parallel. Male inflorescence in leaf axils toward base of branchlets, a lax pedunculate head; flowers many; perianth 4–7-lobed; stamens 8–16; rudimentary ovary pubescent. Female inflorescences in axils of leaves; flowers usually 2, subtended by a single (3-or)4-parted cupule and surrounded by numerous bracts; styles 3, recurved. Cupules woody, (3 or)4-valved; bracts leaflike or filiform to short, triangular.

Nuts (1 or)2(or 3), ovoid to 3-sided. Germination epigeal; cotyledons plicate.

Ten species: N temperate regions; four species (three endemic) in China.

The status of Fagus chienii W. C. Cheng (Contr. Biol. Lab. Chin. Assoc. Advancem. Sci., Sect. Bot. 10: 70. 1935) is uncertain. It is only known

from the type collection (Pingwu, Sichuan) and is similar to F. lucida, except for having longer recurved cupule bracts.

Important timber trees.

1a. Cupules covered with leaflike bracts; leaf blade secondary veins curving upward, fusing near margin 1. F. engleriana

1 Herbarium, Department of Taxonomy, South China Institute of Botany, Chinese Academy of Sciences, Wushan, Guangzhou, Guangdong 510650, People’s

Republic of China.

2 Herbarium, Laboratory of Phytotaxonomy, Fujian Institute of Subtropical Botany, Xiamen, Fujian 361006, People’s Republic of China.

3 Botany Department, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118-4599, U.S.A.

1b. Cupules covered with filiform and recurved or short, triangular bracts; leaf blade secondary veins ending in teeth.

2a. Cupules 2–2.5 cm ................................................................................................................ 2. F. longipetiolata 2b. Cupules 0.7–1.5 cm.

3a. Cupules 0.7–1 cm; leaf blade abaxially with glandular dots and tufts of hairs in axil; cupule bracts linear, recurved ............................................................................................................. 3. F. hayatae 3b. Cupules 1–1.5 cm; leaf blade abaxially without glandular dots and tufts of hairs but with silky pubescence

on midvein; cupule bracts tuberculate, closely appressed .........................................................4. F. lucida

1. Fagus engleriana Seemen in Diels, Bot. Jahrb.Syst. 29: 285. 1900.

米心水青冈 mi xin shui qing gang

Fagus sylvatica Linnaeus var. chinensis Franchet. Trees to 25 m tall. Petiole 0.5–1.5 cm, glabrous; leaf blade ovate, elliptic-ovate, or rarely oblong-ovate, 5–

9(–11) cm, abaxially glaucescent and glabrous except for long silky hairs along veins, base broadly cuneate to sometimes rounded or subcordate, margin sinuate, apex shortly acuminate; secondary veins 9–14 on each side

of midvein, curving upward, fusing near margin. Peduncle 2–7 cm. Cupule 1.5–1.8 cm; basal bracts greenish, leaflike, glabrous, veined; apical bracts brownish, filiform, hairy. Nuts slightly exserted, apex with 3 small wings. Fl. Apr–May, fr. Aug–Oct.

? Broad-leaved and mixed forest on mountain slopes; 1500–2500 m. Anhui, N Guangxi, S Guizhou, Henan, NW Hubei, Hunan, Shaanxi,

E Sichuan, Yunnan, Zhejiang.

2. Fagus longipetiolata Seemen, Bot. Jahrb. Syst. 23(Beibl.

57): 56. 1897.

水青冈 shui qing gang

Fagus bijiensis C. F. Wei & Y. T. Chang; F. brevipetio-lata Hu; F. clavata Y. T. Chang; F. longipes (Oliver) H. Léveillé; F. longipetiolata f. clavata (Y. T. Chang) Y. T. Chang; F. sylvatica Linnaeus var. bracteolis Oliver; F. sylvatica var. longipes Oliver; F. tientaiensis Liou. Trees to 25 m tall. Winter buds to 2 cm. Petiole 1–3.5 cm; leaf blade ovate to ovate-oblong,9–15 cm, abaxially finely densely pubescent and glaucescent, base broadly cuneate to nearly rounded, margin remotely serrate, apex acute to acuminate; secondary veins 9–15 on each side of midvein, ending in teeth. Peduncle 1–10 cm. Cupule 2–2.5 cm; bracts linear, re-curved, pubescent, apical ones to 7 mm but basally shorter. Nut as long or slightly shorter than cupule, with narrowed wings near apex. Fl. Apr–May, fr. Aug–Oct. Broad-leaved evergreen and mixed mesophytic forests on mountain slopes; 300–2400 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang [Vietnam].

3. Fagus hayatae Palibin in Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30(1): 286. 1911. 台湾水青冈 tai wan shui qing gang

Fagus hayatae var. zhejiangensis M. C. Liu & M. H. Wu ex Y. T. Chang & C. C. Huang; F. pashanica C. C. Yang.

Trees to 20 m tall. Winter buds to 1.5 cm. Leaf blade rhomboid-ovate, 3–7 cm, covered with silky pubescence when young, glabrescent except for glandular dots and tufts of hair on midvein and abaxially on axils of secondary veins, base broadly cuneate to nearly rounded, apex acute to shortly acuminate; veins blackish when dry; midvein flexuous toward apex; secondary veins 5–9 on each side of midvein, ending in teeth. Peduncle 0.5–2 cm, pilose. Cupule 7–10 mm; bracts linear, recurved, 1–3 mm, pilose. Nut as long as cupule, with very small wings near apex. Fl. Apr–May, fr. Aug–Oct.

? Mountain ridges and summits in deciduous forests; 1300–2300 m. Hubei, Hunan, Shaanxi, Sichuan, N Taiwan, Zhejiang.

Occurs in three disjunct areas; the plants from each area are slightly different and have been treated as taxonomically distinct.

4. Fagus lucida Rehder & E. H. Wilson in Sargent, Pl. Wilson. 3: 191. 1916.

光叶水青冈 guang ye shui qing gang

Fagus lucida var. opienica Y. T. Chang; F. nayonica Y. T. Chang.

Trees to 25 m tall. Winter buds ca. 1.5 cm. Petiole 0.6–2 cm; leaf blade ovate to elliptic-ovate, 5–11 cm, lustrous green and glabrous except for silky pubescence abaxially on midvein, base broadly cuneate to rounded, margin slightly sinuate, apex acute to shortly acuminate; secondary veins 8–12 on each side of midvein, ending

in minute teeth. Peduncle 0.5–1.5 cm, glabrous. Cupule 1–1.5 cm; bracts tuberculate, closely appressed, triangular mucronate, 1–2 mm, rarely apical ones slightly ascending. Nut slightly exserted, with minute or hardly evident wings near apex. Fl. Apr–May, fr. Sep–Oct.

? Mixed mesophytic forests on mountain slopes; 800–2000 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Zhejiang.

2. CASTANEA Miller, Gard. Dict., Abr. ed. 4, 1: [278]. 1754.

栗属 li shu

Trees or rarely shrubs, deciduous. Bark furrowed. Axillary buds of most apical leaves in false-terminal buds, usually enclosed by 2 outer scales, imbricate within. Leaves spirally arranged but twisted and appearing 2-ranked; secondary

veins ± parallel. Male inflorescences erect catkins; flowers in clusters of 1–3(–5) with each cluster subtended by a bract; perianth 6-parted; stamens 10–12(–20); rudimentary pistil pubescent. Female flowers borne on proximal part of androgynous inflorescences, rarely on a separate inflorescence, usually 3 and subtended by 1 symmetric cupule; ovary 6–9-loculed; styles 6–9; stigmas terminal, minutely punctiform. Cupules splitting into 2–4 valves; bracts spinelike. Nuts 1–3 per cupule. Germination hypogeal.

About 12 species: Asia, Europe, North America; four species (two endemic, one introduced) in China.

Most species are cultivated for edible nuts and durable wood.

1a. Nut 1 per cupule, usually longer than wide ........................................................................................... 4. C. henryi 1b. Nuts usually 2 or 3 per cupule, usually shorter than wide.

2a. Leaf blade abaxially often without scalelike glands but at least when young densely covered with tomentose

or stellate hairs ......................................................................................................................... 1. C. mollissima 2b. Leaf blade abaxially covered with scalelike glands.

3a. Leaf blade glabrous or abaxially sparsely pubescent along veins ........................................... 2. C. seguinii 3b. Leaf blade abaxially tomentose ............................................................................................... 3. C. crenata

1. Castanea mollissima Blume, Mus. Bot. 1: 286. 1850.

栗 li

Castanea bungeana Blume; C. duclouxii Dode; C. fargesii Dode; C. formosana (Hayata) Hayata; C. hupehensis Dode; C. mollissima var. pendula X. Y. Zhou & Z. D. Zhou; C. sativa Miller var. formosana Hayata;C. sativa var. mollissima (Blume) Pampanini; C. vulgaris Lamarck var. yunnanensis Franchet.

Trees to 20 m tall. Branchlets with short pubescence, often also with long spreading hairs. Petiole 1–2 cm; leaf blade elliptic-oblong to oblong-lanceolate, 10–17 cm or rarely shorter, at least along veins abaxially tomentose to softly pubescent, adaxially scalelike glands sometimes absent, base rounded to truncate, margin coarsely serrate, apex acute to acuminate. Male inflorescences 10–20 cm. Cupule densely covered with pubescent spinelike bracts. Nuts usually 2 or 3 per cupule, 2–3 cm in diam. or rarely narrower. Fl. Apr–Jun, fr. Aug–Oct.

Cultivated or wild particularly on mountain slopes; near sea level to 2800 m. Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Liaoning, Nei Mongol, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Xizang, Yun-nan, Zhejiang [Korea].

Extensively cultivated for its edible nuts. Most collections are im-possible to determine if they are cultivated, escaped, or native.

2. Castanea seguinii Dode, Bull. Soc. Dendrol. France 8: 152. 1908.

茅栗 mao li

Castanea davidii Dode.

Trees or shrubs small, rarely to 12 m tall. Stipules nar-rowly lanceolate, 0.7–1.5 cm, deciduous in fruit. Petiole 0.5–1.5 cm; leaf blade oblong-obovate to elliptic-oblong, 6–14 cm, abaxially covered with yellowish brown or grayish, scalelike glands and sparsely hairy along veins when young, base rounded to sometimes subcordate but cuneate when young, margin coarsely serrate, apex acuminate. Male inflorescences 5–12 cm. Female flowers solitary or few per cupule. Cupule 3–5 cm in diam., covered with sparsely pilose spinelike bracts 6–10 mm. Nuts 2 or 3 or rarely more per cupule, 1.5–2 cm in diam. Fl. May–Jul, fr. Sep–Nov. ? Mixed mesophytic forests, thickets, and in orchards; 400–2000 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shanxi, Sichuan, Yunnan, Zhejiang. Cultivated for its edible nuts but not as extensively as Castanea mollissima.

3. Castanea crenata Siebold & Zuccarini, Abh. Math.-Phys. Cl. K?nigl. Bayer. Akad. Wiss. 4: 22

4. 1846.

日本栗 ri ben li

Castanea japonica Blume; C. stricta Siebold & Zuccarini.

Trees or shrubs to 15 m tall. Petiole 0.5–2.5 cm; leaf blade oblong-lanceolate, 8–19 cm, abaxially tomentose and covered with yellowish brown to yellowish gray scalelike glands, base broadly cuneate, rounded, or rarely subcordate, margin finely serrate, apex acuminate to acute; secondary veins ending on teeth and often reduced to long bristlelike points. Male inflorescences 7–20 cm. Female flowers 3 per cupule. Cupule 5–6 cm in diam., covered with spinelike bracts 1–1.5 cm. Nuts 2 or 3 or rarely more per cupule, 2–3 cm in diam., apex sparsely covered with appressed pubescence. Fl. Apr–Jun, fr. Sep–Oct.

Cultivated. Jiangxi (Lu Shan), Liaoning, Shandong (Qingdao Shi), Taiwan [native to Japan, S Korea].

Castanea crenata was introduced from Korea about 1910; var. dulcis Nakai has been reported from NE China, but probably only var. crenata is grown.

4. Castanea henryi (Skan) Rehder & E. H. Wilson in Sargent, Pl. Wilson. 3: 196. 1916.

锥栗 zhui li

Castanopsis henryi Skan in F. B. Forbes & Hemsley, J. Linn. Soc., Bot. 26: 523. 1899; Castanea sativa Miller var. acuminatissima Seemen; C. vilmoriniana Dode. Trees to 30 m tall. Petiole 1–2.5 cm; leaf blade oblong-ovate, oblong-lanceolate, or lanceolate, 10–23 cm, abaxially covered with yellowish brown scalelike glands and sparsely pilose along veins when young, glabrescent, base rounded to broadly cuneate but

narrowly cuneate when young, margin with bristlelike teeth 2–4 mm, apex long acuminate. Male inflores-cences 5–16 cm. Female flowers 1(–3) per cupule. Cupules on a short spike, 2.5–3.5 cm in diam. including bracts, covered with slightly pubescent spinelike bracts. Nut 1 per cupule, globose-ovoid, 1.5–2 cm, longer than wide. Fl. May–Jul, fr. Sep–Oct. ? Mixed mesophytic forests on mountain slopes; 100–1800 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang.

Castanea henryi var. omeiensis W. P. Fang (Acta Phytotax. Sin. 9: 307. 1964) has stellate tomentose hairs on the young leaf blades and 1–3 nuts per cupule, and is probably not a variety of C. henryi but rather a hybrid between C. henryi and C. mollissima.

3. CASTANOPSIS (D. Don) Spach, Hist. Nat. Vég. 11: 142, 185. 1841, nom. cons.

锥属 zhui shu

Quercus Linnaeus [unranked] Castanopsis D. Don, Prodr. Fl. Nepal. 56. 1825; Limlia Masamune & Tomiya; Pasaniopsis Kudo; Shiia Makino.

Trees evergreen. Winter buds ovoid to ellipsoid, with decussate scales. Stipules extrapetiolar. Leaves alternate, distichous, or for a few species spirally arranged. Inflorescences usually unisexual, erect, spicate or paniculate. Male flowers in fascicles of 3–7, rarely solitary and scattered; perianth 5- or 6(–8)-lobed; stamens (8 or)9–12; rudimentary pistil very small, densely covered with curved woolly hairs. Female flowers solitary or in clusters of 3–5(–7) per cupule; staminodes when present opposite perianth lobes; ovary 3-loculed; styles (2 or)3(or 4); stigmas punctiform

or shallow terminal pores. Cupules solitary on rachis, actinomorphic or zygomorphic, rarely indehiscent, completely

or partially enclosing nut; bracts sparsely to densely covering outside of cupule, spinelike or rarely scalelike or tubercles (cupule measurement always includes bracts). Nuts 1–3 per cupule, maturing after 2nd year or rarely in 1st year; abortive ovule apical. Germination hypogeal; cotyledons slightly convex, rarely cerebriform rugose.

About 120 species: tropical and subtropical Asia; 58 species (30 endemic) in China.

Most species are used for timber, and the nuts of many are edible.

1a. Cupules each with 2 or 3(–7) female flowers at anthesis (occasionally on a given inflorescence a cupule can be found with only 1 flower); mature cupules with (1 or)2 or 3 nuts.

2a. Cupule bracts 1–2 cm; leaf blade midvein adaxially slightly raised.

3a. First-year branchlets and leaf blades densely pilose; leaf blade secondary veins 16–22 on each side of

midvein ..................................................................................................................................... 49. C. tessellata 3b. First-year branchlets and leaf blades with scalelike trichomes; leaf blade secondary veins 10–17 on each

side of midvein.

4a. Leaf blade margin serrate; secondary veins 10–13 on each side of midvein ...................... 47. C. xichouensis 4b. Leaf blade margin entire; secondary veins 14–17 on each side of midvein ................................ 48. C. rockii

2b. Cupule bracts less than 1 cm (to 1.5 cm in C. lamontii); leaf blade midvein adaxially impressed or flat.

5a. First-year branchlets and leaf blades abaxially at least adjacent to midvein pilose or villous.

6a. Leaf blade 16–30 × 5–8 cm; secondary veins 16–22 on each side of midvein .............. 50. C. kweichowensis 6b. Leaf blade 10–18 × 2–5 cm; secondary veins 13–17 ......................................................51. C. ceratacantha 5b. First-year branchlets and leaf blades glabrous or apex of young branchlets and midvein of young leaf

blades abaxially with sparse coarse hairs.

7a. Nuts glabrous or with sparse short hairs only around scar when young ....................................... 52. C. fabri 7b. Nuts covered with appressed hairs.

8a. Leaf blade apex mucronate or caudate.

9a. Leaf blade apex mucronate; cupule bracts transversely united to discontinuous cristate rings 55. C. crassifolia 9b. Leaf blade apex caudate and bent; cupule bracts in bundles ................................................ 56. C. chunii

8b. Leaf blade apex acute to acuminate.

10a. Petiole 1.5–3 cm .............................................................................................................. 54. C. lamontii

10b. Petiole 0.8–1.5 cm.

11a. First-year leaf blades glabrous ................................................................................ 57. C. orthacantha

11b. First-year leaf blades abaxially covered with reddish brown to yellowish brown slightly

loose waxy scalelike trichomes.

12a. Leaf blade secondary veins 13–19 on each side of midvein .......................................... 53. C. wattii

12b. Leaf blade secondary veins 9–13 on each side of midvein ................................ 58. C. platyacantha

1b. Cupules each with 1 female flower at anthesis (on a given inflorescence occasionally a cupule can be found with up to 3 flowers); mature cupules with 1(–3) nut.

13a. Cupule bracts scalelike or only bract base transversely adnate to rib rings; nuts maturing and falling in 1st year but empty cupules persisting on rachis; shoot leaves spirally arranged (distichous in C.

uraiana).

14a. Cotyledons cerebriform rugose; young leaf blades glaucous, covered with brown to reddish brown waxy scalelike trichomes.

15a. Base of cupules shortly stipitate; branchlets pubescent ........................................................ 3. C. cerebrina 15b. Base of cupules sessile; branchlets glabrous.

16a. Cupules cupular, enclosing 1/2–2/3 of nut; nut 0.3–0.6 cm in diam. ........................... 1. C. calathiformis 16b. Cupules ellipsoid to ovoid, completely or almost completely enclosing nut; nut 1.1–1.6 cm in diam. ........................................................................................................................................... 2. C. fissa 14b. Cotyledons plano-convex; young leaf blades not glaucous, without or only abaxially with ± adherent waxy scalelike trichomes.

17a. Cupules globose to subglobose, completely or almost completely enclosing nut; nut scar 7–9 mm in

diam. ................................................................................................................................. 6. C. sclerophylla 17b. Cupules shallowly cupular, rarely enclosing more than 1/4–1/3 of nut; nut scar 4–6 mm in diam.

18a. Leaf blade base inaequilateral; fruit maturing on previous year’s branches ......................... 4. C. uraiana 18b. Leaf blade base attenuate; fruit maturing on current year’s branches ............................ 5. C. longzhouica 13b. Cupule bracts spiny or rarely tubercular; nuts maturing and falling in 2nd year, cupules falling with nut; shoot leaves usually distichous.

19a. Cupule outside wall completely covered by spinelike bracts.

20a. Leaf blade margin serrate.

21a. First-year branchlets, petioles, and leaf blades abaxially glabrous or sparsely pubescent.

22a. Leaf blade margin shallowly serrate from middle to apex; secondary veins usually not

reaching margin; petiole rarely longer than 1 cm ................................................................ 8. C. hystrix 22b. Leaf blade margin serrate from near base to apex; secondary veins reaching margin; petiole 1–3 cm.

23a. Leaf blade usually narrower than 5 cm; young leaf blades abaxially covered with

sometimes early glabrescent brownish powdery waxy scalelike trichomes; cupules 2.5–4

cm in diam. ............................................................................................................... 17. C. choboensis 23b. Leaf blade 5–10 cm wide; young leaf blades abaxially thinly covered with reddish brown to yellowish brown waxy scalelike trichomes; cupules 6–8 cm in diam. ......................... 13. C. tibetana 21b. First-year branchlets, petioles, and leaf blades abaxially puberulent.

24a. Transverse section of cupule bracts triangular or trapeziform .......................................... 16. C. clarkei 24b. Transverse section of cupule bracts circular.

25a. Leaf blade secondary veins 15–25 on each side of midvein, reticulate veins abaxially

conspicuous; petiole usually less than 1 cm .................................................................... 14. C. indica 25b. Leaf blade secondary veins 10–15 on each side of midvein, reticulate veins abaxially very slender and evident to inconspicuous; petiole 1–1.8 cm ......................................... 15. C. hainanensis 20b. Leaf blade margin entire or rarely a few leaves shallowly serrate toward leaf blade apex.

26a. Branches and leaf blades glabrous or leaves abaxially with membranous scalelike trichomes.

27a. Cupules 6–8 cm in diam., wall ca. 3 mm thick; bracts 2–3 cm; nuts oblate, puberulent, scar covering

1/3 of nut ....................................................................................................................... 7. C. kawakamii 27b. Cupules 2–3 cm in diam., wall ca. 1 mm thick; bracts 0.4–1 cm; nuts broadly conical,

glabrous, scar covering only basal part of nut ...................................................................... 34. C. eyrei 26b. Branches and leaf blades hairy (at least on 1st-year branchlets and abaxial midvein) or leaf blades abaxially covered with waxy scalelike trichomes.

28a. Leaf blade base cordate, shallowly auriculate, or rarely rounded; petiole 1–3(–5) mm ...... 12. C. fordii 28b. Leaf blade base not cordate or auriculate but may be rounded; petiole (4–)8–20 mm.

29a. Leaf blade 4–10 cm.

30a. Infructescences ca. 15 cm; cupules 2.5–4 cm in diam., bracts 0.6–1 cm ....................... 8. C. hystrix

30b. Infructescences 4–8 cm; cupules 5–6 cm in diam., bracts 1–2 cm ............................. 9. C. concinna 29b. Leaf blade 10–25 cm.

31a. Petiole 1–2 cm; leaf blade 15–25 cm, midvein adaxially usually impressed, secondary

veins

12–16 on each side of midvein; nuts oblate, 1.5–2 cm in diam., scar covering ca. 1/3 of

nut

............................................................................................................................. 10. C. mekongensis 31b. Petiole 0.8–1 cm; leaf blade 10–15 cm, midvein adaxially raised, secondary veins 9–12

on

each side of midvein; nuts broadly conical, 2–3.5 cm in diam., scar covering only basal

part

of nut ................................................................................................................ 11. C. globigemmata

19b. Cupule outside wall visible (if spinelike bracts completely cover wall then spines transversely adnate

to cristate rings)

32a. Scar covering basal 2/3 or more of nut.

33a. Rachis of infructescences covered with pale grayish brown, feltlike, short hairs, glabrescent;

cupule

bracts 2–6 mm ............................................................................................................ 19. C. argyrophylla 33b. Rachis of infructescences glabrous or very shortly and sparsely mealy puberulent; cupule

bracts

7–12 mm ..................................................................................................................... 20. C. tcheponensis

32b. Scar at base or covering only basal 1/3 of nut.

34a. Mature nuts with appressed hairs.

35a. Rachis of infructescences 5–8 mm thick and leaf blade margin entire.

36a. Cupules 2–2.5 cm in diam.; petiole 0.3–0.6 cm; leaf blade margin subconvolute 23. C. damingshanensis

36b. Cupules 2.5–4.5 cm in diam.; petiole 1–3 cm; leaf blade margin not subconvolute.

37a. Leaf blade 26–45 cm, young leaf blades abaxially with slender stellate hairs along

midvein;

secondary veins 16–20 or more on each side of midvein; petiole 2–3 cm ............ 21. C. megaphylla 37b. Leaf blade 8–15 cm, young leaf blades abaxially covered with yellowish brown tightly

adnate

layers of waxy scalelike trichomes; secondary veins 10–13 on each side of midvein;

petiole

1–2 cm ................................................................................................................... 22. C. nigrescens 35b. Rachis of infructescences less than 5 mm thick but if 5 mm thick then leaf blade margin with short

and sharp serrations.

38a. Petiole 2.5–3.5 cm; young leaf blades abaxially sometimes with slender stellate hairs 24. C. jianfenglingensis 38b. Petiole rarely to 2.5 cm; young leaf blades abaxially without stellate hairs.

39a. Leaf blade secondary veins and midvein adaxially slightly raised; cupules 3–3.5 cm

in diam. ....................................................................................................... 25. C. remotidenticulata 39b. Leaf blade secondary veins and sometimes midvein adaxially impressed; cupules 1.5–3

cm in diam.

40a. Leaf blade margin mostly entire.

41a. Leaf blade lanceolate, 1.5–4 cm wide, apex caudate to sometimes shortly acute,

secondary

veins 9–13 on each side of midvein; nuts densely pilose ................................ 26. C. tonkinensis 41b. Leaf blade ovate-elliptic, ovate, or oblong, 4–6 cm wide, apex acuminate, secondary

veins

13–17 on each side of midvein; nuts puberulent ...................................................... 28. C. boisii 40b. Leaf blade margin serrate but some leaves on same plant may be entire.

42a. First-year leaf blades concolorous or adaxially darker green and abaxially greenish 29. C. densispinosa

42b. First-year leaf blades not concolorous.

43a. Cupules ca. 3 cm in diam.; leaf blade midvein adaxially usually slightly raised at

base ...................................................................................................................32. C. oblonga 43b. Cupules 1.5–2 cm in diam.; leaf blade midvein adaxially impressed.

44a. Petiole usually shorter than 1 cm; leaf blade abaxially covered with slightly

adherent

waxy scalelike trichomes .........................................................................30. C. ledongensis 44b. Petiole ca. 1.5 cm; leaf blade abaxially covered with early glabrescent waxy

scalelike trichomes ..................................................................................31. C. subuliformis

34b. Mature nuts glabrous or glabrescent, or sparsely covered with appressed hairs only near apex, or rarely puberulent.

45a. Cupules 0.7–2(–2.2) cm in diam.; bracts 1–6 mm.

46a. Leaf blade margin predominantly entire but sometimes shallowly serrate at apex.

47a. First-year leaf blades abaxially greenish; cupules 0.7–1.1 cm in diam.; leaf blade

secondary veins

7–10 on each side of midvein ....................................................................................... 40. C. fleuryi 47b. First-year leaf blades abaxially red-brown; cupules 1.2–2.2 cm in diam.; leaf blade

secondary veins 11–18 on each side of midvein.

48a. Petiole shorter than 1 cm; leaf blade secondary veins 14–18 on each side of midvein;

leaf blades abaxially and infructescence rachises densely covered with early glabrescent

powdery waxy scalelike trichomes ............................................................... 41. C. rufotomentosa 48b. Petiole 1–1.5 cm; leaf blade secondary veins 11–14 on each side of midvein; leaf blades abaxially and infructescence rachises pubescent, covered with slightly adherent waxy

scalelike trichomes .............................................................................................. 42. C. tribuloides 46b. Leaf blade margin predominantly serrate but sometimes entire.

49a. First-year leaf blades abaxially with a thin greenish layer of adnate scalelike trichomes when young, concolorous with age ................................................................. 43. C. wenchangensis 49b. First-year leaf blades abaxially covered with reddish brown to yellowish brown waxy scalelike trichomes, remaining distinctly different in color from adaxial surface with age.

50a. Leaf blade apex acute to rounded; midvein adaxially slightly raised; leaf blade widest usually from middle to apex ................................................................................... 44. C. delavayi 50b. Leaf blade apex acuminate to caudate; midvein adaxially impressed when dry (flat

when fresh); leaf blade widest from base to middle.

51a. Petiole base not pillow-shaped; mature cupules rarely splitting; nut scar 8–9 mm in

diam. ................................................................................................................ 45. C. echinocarpa 51b. Petiole base pillow-shaped; mature cupules splitting at apex; nut scar ca. 5 mm in diam.

46. C. carlesii

45b. Cupules 2–3.5 cm in diam.; bracts 4–15 mm.

52a. Leaf blade midvein adaxially raised at least from base to middle of blade.

53a. Petiole 1.5–2 cm; rachis of infructescences pubescent; mature cupules and bracts

glabrescent ............................................................................................................... 33. C. chinensis 53b. Petiole 0.7–1.2 cm; rachis of infructescences glabrous; mature cupules and bracts

pubescent.

54a. Bracts usually not in bundles; nuts broadly conical ..................................................... 34. C. eyrei

54b. Bracts usually in bundles; nuts subglobose but apex pointed ...................................... 35. C. ferox 52b. Leaf blade midvein adaxially flat to impressed.

55a. Leaf blade margin predominantly serrate.

56a. Cupule splitting irregularly ...................................................................................... 36. C. fargesii

56b. Cupule splitting regularly.

57a. Leaf blade 4–8 cm wide; bud scales and 1st-year branchlets covered with reddish

brown

slightly loose waxy scalelike trichomes; petiole 1–2.5 cm; cupules 2.5–3.5 cm in diam.

.............................................................................................................................. 38. C. jucunda 57b. Leaf blade 1.5–3.5 cm wide; bud scales and 1st-year branchlets glabrous or covered

with

early glabrescent waxy scalelike trichomes; petiole 0.3–0.7 cm; cupules 2–2.2 cm in

diam. ................................................................................................................ 39. C. hupehensis 55b. Leaf blade margin predominantly entire.

58a. Leaf blade apex caudate ......................................................................................... 27. C. amabilis

58b. Leaf blade apex acute to acuminate.

59a. Petiole 2–2.5 cm; leaf blade elliptic to obovate-elliptic; cupule bracts 0.4–0.8 cm 37. C. ouonbiensis

59b. Petiole 1–2 cm; leaf blade oblong, obovate-oblong, lanceolate, or rarely ovate; cupule bracts 0.8–1.5 cm.

60a. Young shoots tufted puberulent; leaf blade abaxially stellate cespitose along midvein;

cupule bracts 1.3–1.5 cm; nuts ellipsoid .......................................................18. C. longispina 60b. Young shoots from middle to apex with scalelike trichomes; leaf blade abaxially with a

thick and mealy layer of scalelike trichomes; cupule bracts 0.8–1 cm; nuts conical to

subglobose .......................................................................................................... 36. C. fargesii

1. Castanopsis calathiformis (Skan) Rehder & E. H. Wilson in Sargent, Pl. Wilson. 3: 204. 1916.

瘰丝锥 bao si zhui

Quercus calathiformis Skan in F. B. Forbes & Hemsley, J. Linn. Soc., Bot. 26: 508. 1899; Lithocarpus calathiformis (Skan) A. Camus; Pasania calathiformis (Skan) Hickel & A. Camus; Synaedrys calathiformis (Skan) Koidzumi.

Trees 5–10(–20) m tall. First-year branchlets black, dark black-brown, or dusky when dry, glabrous. Leaves spirally arranged, yellowish brown to gray; petiole 1–2.5 cm; leaf blade oblong to obovate-elliptic, very variable in size, often 15–25 × 5–9 cm, thickly papery, covered with brown-red slightly adnate small lamellate waxy scalelike trichomes when young, base cuneate, margin from base to middle undulate and crenate, apex acute, acuminate, or rounded; midvein adaxially raised; secondary veins 20–28 on each side of midvein or rarely fewer. Female inflorescences usually borne more toward apex of branches and if axillary then solitary, 8–16 cm. Infructescences 10–20 cm. Cupules yellowish brown to dark grayish brown when dry, cupular, 0.8–1.2 cm in diam., wall ca. 1 cm thick; bracts scalelike, triangular, annular or sometimes imbricate when young, thickened with age, adnate, arranged in 4–7 rib rings, waxy. Nut ovoid to ellipsoid, 1–1.5 cm × 3–6 mm, apically brown-red tomentose; scar basal, 3–6 mm in diam. Fl. Mar–May, fr. Oct–Dec.

Mixed and broad-leaved evergreen forests; 700–2200 m. SE Xizang, S Yunnan [Laos, Myanmar, N Thailand, N Vietnam].

Castanopsis calathiformis,C. cerebrina,C. fissa,C. longzhouica,C. sclerophylla, and C. uraiana form a group of related species with 1 flower per cupule, and cupule bracts reduced to scales. The first two species are also united by having plicate cotyledons, and the latter three by having flat-convex cotyledons.

2. Castanopsis fissa (Champion ex Bentham) Rehder & E. H. Wilson in Sargent, Pl. Wilson. 3: 20

3. 1916.

黧蒴锥 li shuo zhui

Quercus fissa Champion ex Bentham, Hooker’s J. Bot. Kew Gard. Misc. 6: 114. 1854; Castanopsis fissoides Chun & C. C. Huang ex Luong; C. tunkinensis (Drake) Barnett (1944), not C. tonkinensis Seemen (1897); Lithocarpus fissus (Champion ex Bentham) A. Camus; Pasania fissa (Champion ex Bentham) Oersted; Q. tunkinensis Drake; Shiia fissa (Champion ex Bentham) Kudo; Synaedrys fissa (Champion ex Bentham) Koidzumi. Trees; bud scales, shoot apexes, and young leaf blades abaxially yellowish brown puberulent and covered with rust-colored, small, lamellate, slightly adnate, waxy scalelike trichomes. Branchlets red-purple, prominently angular. Leaves spirally arranged, yellowish brown to gray; petiole 1–2.5 cm; leaf blade oblong to obovate-elliptic, very variable in sizes, often 15–25 × 5–9 cm, thickly papery, base cuneate, margin from base to middle undulate and crenate, apex acute, acuminate, or rounded; midvein raised adaxially; secondary veins 20–28 on each side of midvein or rarely fewer. Infructescence 8–18 cm; rachis glabrous. Cupule 1–1.5 cm in diameter, irregularly 2- or 3(or 4)-valved with valves often curled, covered with dark reddish brown mealy waxy scalelike trichomes, apex slightly cuspidate, wall 0.5–1 mm thick; bracts scalelike, triangular to squarish, imbricate when young but mostly united and in ring ribs with age. Nut globose to elliptic, 1.3–1.8 × 1.1–1.6 cm, apically brown-red tomentose; scar basal,

4–7 mm in diam. Fl. Apr–Jun, fr. Oct–Dec.

Broad-leaved evergreen forests; below 1600 m. S Fujian, Guangdong, Guangxi, S Guizhou, Hainan, S Hunan, S Jiangxi, SE Yunnan [N Thailand, N Vietnam].

Cupule shape of this species ranges from ellipsoid to ovoid and correlates with the distribution. Ellipsoid cupules occur in Fujian, E Guangdong, Jiangxi, and Hunan, while ovoid cupules occur in more western regions such as SE Yunnan and neighboring regions, including W Hainan and NE Vietnam.

3. Castanopsis cerebrina (Hickel & A. Camus) Barnett, Trans. & Proc. Bot. Soc. Edinburgh 34: 183. 194

4.

毛叶杯锥 mao ye bei zhui

Pasania cerebrina Hickel & A. Camus, Ann. Sci. Nat., Bot., sér. 10, 3: 408. 1921; Lithocarpus cerebrinus (Hickel & A. Camus) A. Camus.

Trees; young branchlets and leaf blades reddish brown, densely puberulent with simple and stellate hairs and covered with slightly loose, small, lamellate, waxy scalelike trichomes. Branchlets red-purple, prominently angular. Leaves spirally arranged, yellowish brown to gray; petiole 1–2.5 cm; leaf blade oblong to obovate-elliptic, often 15–25 × 5–9 cm, thickly papery, base cuneate, margin from base to middle undulate and crenate, apex acute, acuminate, or rounded; midvein adaxially raised; secondary veins 20–28 on each side of midvein or rarely fewer. Rachis of female inflorescences sparsely puberulent and covered with

waxy scalelike trichomes. Cupule cupular, 1.8–2.2 × 1.5–2.5 cm, base stipitate, stipe 5–8 mm, wall 0.5–1 mm thick; bracts mound-shaped, arranged in disconnected rings, sparsely puberulent, covered with waxy scalelike trichomes. Nut narrowly conical, 2–2.5 × 1.4–2.2 cm, tomentulose, apex sometimes 3-angled; scar 0.8–1.2 cm in diam. Fl. Apr, fr. Oct.

Broad-leaved evergreen forests; 200–700 m. SE Yunnan [N Thailand, N Vietnam].

4. Castanopsis uraiana (Hayata) Kanehira & Hatusima, Trans. Nat. Hist. Soc. Taiwan 29: 15

5. 1939.

淋漓锥 lin li zhui

Quercus uraiana Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30(1): 299. 1911; Limlia uraiana (Hayata) Masamune & Tomiya; Lithocarpus uraiana (Hayata) Hayata; Pasania uraiana (Hayata) Schottky; Q. paohangii Chun & Tsiang; Q. randaiensis Hayata; Shiia uraiana (Hayata) Kanehira & Hatusima; Synaedrys uraiana (Hayata) Koidzumi.

Trees. Young shoots often sparsely puberulent and with small, lamellate, waxy scalelike trichomes or glabrous, sparsely lenticellate. Leaves distichous; petiole 0.7–1.5 cm; leaf blade ovate-elliptic, ovate, lanceolate, or sometimes obovate, 7–13 × 2–3 cm, papery, abaxially covered with brown to reddish brown, tight, waxy scalelike trichomes when young but grayish brown to nearly concolorous with age, base attenuate and in-aequilateral, margin apically sparsely serrate or entire, apex slightly pointed, acute, or caudate and often bent

to 1 side; midvein adaxially flat to slightly impressed or from middle to apex slightly raised; secondary veins 7–12 on each side of midvein, very slender, evident. Infructescences 5–10 cm; rachis ca. 2 mm thick, sparsely gray lenticellate. Cupule shallowly cupular, 5–6 × 7–12 mm, enclosing 1/4–1/3 of nut, wall less than 1 mm thick; bracts reduced to scales, triangular to ovate, adnate, imbricate, dusky puberulent. Nut 1 per cupule, broadly conical, 0.7–1.5 cm; scar basal, 5–6 mm in diam. Fl. Mar–May, fr. Sep–Oct of following year.

? Broad-leaved evergreen forests; 400–1500 m. Fujian, E to N Guangdong, NE Guangxi, S Hunan, S Jiangxi, Taiwan.

5. Castanopsis longzhouica C. C. Huang & Y. T. Chang, Guihaia 5: 18

6. 1985.

龙州锥 long zhou zhui

Trees; branchlets, leaf blades, and infructescences gla-brous. Young shoots grayish, sparsely lenticellate; lenticels grayish, slightly raised. Petiole 1–1.5 cm; leaf blade elliptic, ovate-elliptic, or sometimes obovate, 8–10 cm, firmly papery, slightly brittle, concolorous, abaxially with a thin layer of closely adherent grayish waxy scalelike trichomes, base attenuate, margin from middle to apex dentate to crenate, apex acute and slightly pointed; midvein abaxially prominently raised; secondary veins 9–12 on each side of midvein. Female inflorescences 1–2 cm; flowers few, solitary along rachis. Infructescences almost as long as rachis, sparsely lenticellate; lenticels grayish. Cupule shallowly cupular, 5–6 mm, covering base or no more than basal

1/4 of nut; bracts imbricate, adnate, broadly ovate to ovate-elliptic, gray puberulent, margin slightly membranous. Nut broadly conical, 1–1.5 cm in diam.; scar basal, ca. 4 mm in diam. Fl. Feb–Mar, fr. Aug–Sep. ? Broad-leaved evergreen forests; 400–600 m. Guangxi (Longzhou Xian).

6. Castanopsis sclerophylla (Lindley & Paxton) Schottky, Bot. Jahrb. Syst. 47: 638. 1912.

苦槠 ku zhu

Quercus sclerophylla Lindley & Paxton, Paxt. Fl. Gard. 1: 59. 1850; Lithocarpus chinensis (Abel) A. Camus; Q. chinensis Abel (1818), not Castanopsis chinensis Hance (1868); Q. cuspidata Thunberg var. sinensis A. de Candolle; Synaedrys sclerophylla (Lindley & Paxton) Koidzumi.

Trees; branches and leaf blades glabrous. Young shoots reddish brown, slightly angulate. Petiole 1.5–2.5 cm; leaf blade oblong, ovate-elliptic, or obovate-elliptic, 7–15 cm, leathery, adaxially silver-gray with age, base rounded to broadly cuneate and usually inaequilateral, margin from middle to apex serrulate or rarely entire, apex acuminate, cuspidate, or shortly caudate; midvein slightly impressed from base to middle and slightly raised from middle to apex; secondary veins 10–15 on each side of midvein, prominent to very slender, evident. Rachis of inflorescences glabrous. Female inflorescence ca. 15 cm. Infructescences 8–15 cm. Cupule globose to subglobose, 1.2–1.5 cm in diam., completely or almost completely enclosing nut, irregu-larly valved, outside yellowish brown puberulent, wall to 1 mm thick; bracts scalelike, 3- or 4-angled, sometimes only base connate, in annular umbones. Nut 1(–3) per cupule, subglobose, 1–1.4 cm in diam., tomentulose, apex mucronulate; scar basal, 7–9 mm in diam. Fl. Apr–May, fr. Oct–Nov.

? Broad-leaved evergreen forests; 200–1000 m. Anhui, Fujian, Guangxi, NE Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, E Sichuan, Zhejiang.

The nuts are used for food.

Castanopsis ×kuchugouzhui C. C. Huang & Y. T. Chang (Guihaia 16: 301. 1996) is a hybrid between C. sclerophylla and C. tibetana with a population on Yuelu Shan in Changsha Shi, Hunan. The bracts apically and along the sides of the cupule are 5–8 mm and spinelike, like those of C. tibetana, but among them, as well as the basal bracts, are small, triangular to multiangular, and lamellate bracts like those of

C. sclerophylla.

7. Castanopsis kawakamii Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30(1): 300. 1911.

吊皮锥 diao pi zhui

Castanopsis greenii Chun; C. oerstedii Hickel & A. Camus.

Trees. Branches glabrous; young shoots dark reddish brown, glabrous, sparsely lenticellate; lenticels dark gray. Petiole 1–2.5 cm; leaf blade ovate to lanceolate,

6–12 × 2–5 cm, dark reddish brown when young and remaining so abaxially, leathery with age, base broadly cuneate to rounded and inaequilateral, margin entire or rarely 1–3-toothed near apex, apex caudate; midvein adaxially slightly raised from base to middle but flat to slightly impressed from middle to apex; secondary veins 9–12 on each side of midvein; tertiary veins reticulate, conspicuous. Female inflorescences 5–10 cm; rachis glabrous. Cupule globose, 6–8 cm in diam., splitting into 4(or 5) segments when mature, inside densely dusky tomentulose, wall ca. 3 mm thick; bracts spinelike, connected to radial branched bundles in mid-dle part of cupule or slightly basally, 2–3 cm, sparsely pubescent to nearly glabrous. Nut 1 per cupule, oblate, 1.2–1.5 × 1.7–2 cm, densely orangish brown puberulent; scar covering ca. 1/3(–1/2) of nut. Fl. Mar–Apr, fr.

Aug–Oct of following year.

Broad-leaved evergreen forests; below 1000 m. S Fujian, Guangdong, SE Guangxi, S Jiangxi, C Taiwan [Vietnam].

Reports of Castanopsis borneensis King from Taiwan are referable to C. kawakamii. Castanopsis concinna,C. fordii, C. globigemmata,C. hainanensis,C. hystrix,C. indica,C. kawakamii,C. mekongensis, and C. tibetana form a group of related species with cupules splitting regularly into 4 parts, bract with varying length that are basally connate into fascicles, and a scar that occupies the basal 1/4 to 1/3 of the nut.

8. Castanopsis hystrix J. D. Hooker & Thomson ex A. de Candolle, J. Bot. 182. 1863.

红锥 hong zhui

Castanea bodinieri H. Léveillé & Vaniot; Castanopsis bodinieri (H. Léveillé & Vaniot) Koidzumi; C. brunnea (H. Léveillé) A. Camus; C. lohfauensis Hu; C. tapuensis Hu; Quercus brunnea H. Léveillé.

Trees. Young shoots purple-brown, slender, sparsely to densely puberulent and with yellowish brown small lamellate waxy scalelike trichomes. Petiole ca. 1 cm or rarely longer; leaf blade lanceolate to obovate-elliptic,

4–9 × 1.5–4 cm or rarely smaller or larger, papery to thinly leathery, pubescent when young but early glabrescent, at least adaxially along midvein with very lax and thick or tight and thin, reddish brown to yel-lowish brown, small, lamellate, waxy scalelike trichomes, base sharply acute to rounded and inaequilateral, apex mucronate to caudate; midvein adaxially impressed; secondary veins 9–15 on each side of midvein, very slender, evident. Female inflorescence solitary in leaf axil. Infructescence ca. 15 cm. Cupule globose, 2.5–4 cm in diam., splitting into 4 segments, wall ca. 2.5 mm thick; bracts spinelike, completely covering cupule, 6–10 mm, sparsely puberulent, base of some connate into bundles. Nut 1 per cupule, broadly conical, 1–1.5 × 0.8–1.3 cm, glabrous; scar basal. Fl. Apr–Jun, fr. Aug–Nov of following year.

Broad-leaved evergreen forests; near sea level to 1600 m. SE Fujian, Guangdong, Guangxi, Guizhou, Hainan, SW Hunan, SE Xizang

(Mêdog Xian), S Yunnan [Bhutan, Cambodia, NE India, Laos, Myan-mar, Nepal, Sikkim, Vietnam].

9. Castanopsis concinna (Champion ex Bentham) A. de Can-dolle in Hance, J. Bot. 1: 182. 1863.

华南锥 hua nan zhui

Castanea concinna Champion ex Bentham, Hooker’s J. Bot. Kew Gard. Misc. 6: 115. 1854; Castanopsis oblongifolia W. C. Cheng & C. S. Chao.

Trees; young shoots and rachis of inflorescences yellowish brown to reddish brown puberulent and with early glabrescent, very thick, small, lamellate, waxy scalelike trichomes. Petiole ca. 4–12 mm; leaf blade elliptic, oblong-orbicular, or sometimes oblanceolate,

5–10 × 1.5–3.5(–5) cm, leathery, brittle, margin and midvein pilose when young, base shortly attenuate to broadly cuneate and usually symmetric but sometimes slightly asymmetric, margin entire or apically sparsely serrulate, apex mucronate to acuminate; midvein adaxially prominently impressed; secondary veins 12–16 on each side of midvein. Female inflorescences 5–10 cm. Infructescences 4–8 cm; rachis 4–6 mm thick. Cupule globose, 5–6 cm in diam., splitting into 4 regular segments, wall 2–4 mm thick; bracts spinelike, entirely covering cupule, 1–2 cm, puberulent, base connate into bundles. Nut 1 per cupule, ± conical, ca. 1 × 1.4 cm, densely pubescent; scar covering ca. 1/3 but no more than 1/2 of nut. Fl. Apr–May, fr. Sep–Oct of following year.

? Broad-leaved evergreen forests; below 500 m. S Guangdong, S Guangxi.

10.Castanopsis mekongensis A.Camus,Bull. Soc. Bot. France 85: 653. 1938 [1939].

湄公锥 mei gong zhui

Castanopsis fohaiensis Hu; C. lantsangensis Hu; C. wangii Hu & W. C. Cheng.

Trees; young shoots, petioles, leaf blades abaxially, and rachis of inflorescences densely pubescent. Petiole 1–2 cm; leaf blade ovate-elliptic, broadly elliptic, or sometimes ovate, 15–25 × 5–8 cm, thickly papery to leathery, abaxially grayish, adaxially dark black-brown when young, grayish brown to yellowish gray when dry, puberulent on midvein, base rounded to acute and symmetric or inaequilateral, margin entire, apex acuminate; midvein and secondary veins adaxially usually impressed; secondary veins 12–16 on each side of midvein. Infructescence ca. 10 cm or rarely longer. Cupule globose, 5–6 cm in diam., wall ca. 2 mm thick; bracts spinelike, entirely covering cupule, 1–1.5 cm, sparsely pubescent, base connate into bundles. Nut 1 per cupule, oblate, 1.3–1.6 × 1.5–2 cm, densely pubescent; scar covering ca. 1/3 of nut. Fl. Mar–Apr, fr. Aug–Oct of following year.

Broad-leaved evergreen forests; 600–2000 m. S to SE Yunnan [Laos].

11. Castanopsis globigemmata Chun & C. C. Huang in C. C. Huang & Y. T. Chang, Guihaia 16: 300. 1996.

圆芽锥 yuan ya zhui

Trees. Branchlet black-brown when dry, densely lenticellate; lenticels slightly raised. Petiole 8–10 mm; leaf blade ovate-elliptic to lanceolate, 10–15 × 3.5–5 cm, nearly leathery, pilose and with tight brown, waxy scalelike trichomes, abaxially when young and on both sides along midvein, base broadly cuneate to acute and inaequilateral, margin entire or with 1–3 shallow teeth from middle to apex, apex long acuminate to caudate; midvein at least from base to middle adaxially raised; secondary veins 9–12 on each side of midvein. Rachis of inflorescences sparsely puberulent and with brown, stellate, waxy scalelike trichomes. Infructescences 5–7 cm; cupules clustered from middle to apex of rachis. Cupule globose, 6–7 cm in diam., splitting into 4 regular segments when mature, wall 3–4 mm thick; bracts spinelike, entirely covering cupule, 1–1.5 cm, base connate into bundles. Nut 1 per cupule, broadly conical, 2–3.5 cm in diam., densely pubescent; scar basal. Fl. Aug–Sep, fr. Oct–Nov of following year.

? Broad-leaved evergreen forests; ca. 1400 m. SE Yunnan (Pingbian Miaozu Zizhixian).

12. Castanopsis fordii Hance, J. Bot. 22: 230. 1884.

毛锥 mao zhui

Trees; bud scales, young shoots, petioles, leaf blades abaxially, and rachis of inflorescences densely covered with brown to reddish brown, slightly rough long tomentum. Petiole 1–3(–5) mm, usually tapering toward apex; leaf blade oblong, lanceolate, or oblanceolate-oblong, 9–18 × 3–7 cm, leathery, abaxially reddish brown when young but grayish brown to grayish with age, base cordate, shallowly auriculate, or rarely rounded, margin entire and revolute, apex acute, mucronate, or rarely rounded; midvein adaxially prominently impressed; secondary veins 14–18 on each side of midvein or fewer. Infructescences dense, 6–12 cm. Cupule 5–6 cm in diam., splitting into 4(or 5) regular segments, wall 3–4 mm thick; bracts spinelike, entirely covering cupule, 1–2 cm, pubescent, base connate into many bundles. Nut 1 per cupule, ± conical, 1.2–1.5 × 1.5–2 cm, densely pubescent; scar covering ca. 1/3 of nut. Fl. Mar–Apr, fr. Sep–Oct of following year.

? Broad-leaved evergreen forests; below 1200 m. S Fujian, Guangdong, SE Guangxi, S Hunan, S Jiangxi, S Zhejiang.

13. Castanopsis tibetana Hance, J. Bot. 13: 367. 1875.

钩锥 gou zhui Castanopsis chengfengensis Hu; Quercus franchetiana H. Léveillé ex A. Camus.

Trees. Young shoots black to black-brown when dry, glabrous. Petiole 1.5–3 cm; leaf blade ovate-elliptic, ovate, oblong, or obovate-elliptic, 15–30 × 5–10(–13) cm, leathery, dark purple-brown when young, abaxially thinly covered with reddish brown to yellowish brown, waxy scalelike trichomes, base rounded to shortly cuneate and symmetric or sometimes inaequilateral, margin serrate except basally entire, apex acuminate, mucronate, or caudate; midvein adaxially impressed; secondary veins 15–18 on each side of midvein. Female inflorescences 5–25 cm; rachis 4–6 mm thick. Cupule globose, 6–8 cm in diam., splitting into 4(or 5) regular segments, wall 3–4 mm thick; bracts spinelike, entirely covering cupule, 1.5–2.5 cm, subglabrous to sparsely puberulent, base usually connate into bundles. Nut 1 per cupule, ± conical, 1.5–1.8 × 2–2.8 cm, hairy; scar covering ca. 1/4 of nut. Fl. Apr–May, fr. Aug–Oct of following year.

? Broad-leaved evergreen forests; below 1500 m. S Anhui, Fujian, Guangdong, Guangxi, Guizhou, SW Hubei, Hunan, Jiangxi, SE Yunnan, S Zhejiang.

14. Castanopsis indica (Roxburgh ex Lindley) A. de Candolle in Hance, J. Bot. 1: 182. 1863.

印度锥 yin du zhui

Castanea indica Roxburgh ex Lindley in Wallich, Pl. Asiat. Rar. 2: 5. 1830; Castanopsis macrostachya Hu; C. sinensis A. Chevalier; C. subacuminata Hayata; Quercus indica (Roxburgh ex Lindley) Drake.

Trees; young shoots, petioles, leaf blades abaxially, and rachis of inflorescences yellowish brown puberulent. Petiole 5–10(–15) mm; leaf blade ovate-elliptic, elliptic, or sometimes obovate-elliptic, 9–20 × (4–)6–10 cm, thickly papery, abaxially puberulent or glabrescent, base cuneate to rounded and usually inaequilateral, margin serrate except basally entire, apex mucronate to acuminate; midvein adaxially impressed; secondary veins 15–25 on each side of midvein. Female inflorescence ca. 40 cm. Infructescences dense, 10–27 cm. Cupule globose, 3.5–4 cm in diam., usually splitting into 4 segments when mature, wall ca. 1 mm thick; bracts spinelike, entirely covering cupule, to 1.5 cm, straight or bent, base connate into bundles. Nut 1(or 2) per cupule, broadly conical, 1–1.4 cm in diam., densely hairy; scar covering ca. 1/4 of nut. Fl. Mar–May, fr. Sep–Nov of following year.

Broad-leaved evergreen forests; below 1500 m. S Guangdong, S Guangxi, S Hainan, Taiwan, SE Xizang (Mêdog Xian), S Yunnan [Bangladesh, Bhutan, NE India, Laos, Myanmar, Nepal, Sikkim, Thailand, Vietnam].

15. Castanopsis hainanensis Merrill, Philipp. J. Sci. 21: 340. 1922.

海南锥 hai nan zhui

Trees; branchlets, petioles, young leaf blades abaxially, rachis of inflorescences, and perianth covered with reddish brown, dusky, or grayish brown, very short, felted puberulence. Petiole 1–1.8 cm; leaf blade obovate, obovate-elliptic, ovate-elliptic, or broadly ovate, 5–12(–17) × 2.5–5(–6) cm, thickly papery to nearly leathery, abaxially often grayish with age, base acute to broadly cuneate, margin serrate except basally entire, apex rounded to mucronate; midvein adaxially impressed but often slightly raised on sprouted branch leaves; secondary veins 10–15(–18) on each side of midvein. Infructescences 10–17 cm; rachis 5–6 mm thick. Cupule 4–5 cm in diam., wall ca. 1 mm thick; bracts spinelike, congested, entirely covering cupule, to 1.5 cm, basally connate. Nut 1 per cupule, broadly conical, 1.2–1.5 × 1.6–2 cm, densely pubescent; scar exceeding base of nut. Fl. Mar–Apr, fr. Aug–Oct of following year.

? Broad-leaved evergreen forests; below 400 m. Hainan.

Could be recognized as an insular subspecies of Castanopsis indica. The status of C. undulatifolia G. A. Fu (Guihaia 14: 301. 1994) is uncertain, but it may be conspecific with C. hainanensis.

16. Castanopsis clarkei King ex J. D. Hooker, Fl. Brit. India 5: 623. 1888.

棱刺锥 leng ci zhui

Trees; young shoots and rachis of inflorescences densely puberulent. Winter bud scales velutinous. Petiole 1.5–3 cm; leaf blade elliptic to oblong, 10–20 × 5–9 cm, thickly papery to nearly leathery, base acute and inaequilateral, margin serrate except basally entire, apex mucronate; midvein abaxially conspicuously raised, adaxially slightly impressed; secondary veins 14–20. Female inflorescences ca. 20 cm. Infructescence rachis 4–5 mm thick. Cupule subglobose, 3.5–4(–5) cm in diam., wall ca. 1.5 mm thick; bracts spinelike, congested, entirely covering cupule, 1–1.5 cm, 3- or 4-angled in cross section, free or base connate into bundles, nearly glabrous. Nut 1 per cupule, broadly conical, 1.4–1.6 cm, apex cuspidate; scar basal. Fl.

Mar–May, fr. Oct–Dec of following year.

Broad-leaved evergreen forests; 500–800 m. SE Xizang (Mêdog Xian), S Yunnan [NE India, NE Myanmar].

17. Castanopsis choboensis Hickel & A. Camus, Notul. Syst. (Paris) 4: 122. 1928.

窄叶锥 zhai ye zhui

Trees. Petiole 1–2 cm; leaf blade narrowly elliptic to lanceolate, 8–15 × 3–5 cm or rarely larger, thickly papery, base acute to cuneate and often oblique, margin serrate-dentate, apex mucronate to caudate; midvein slightly impressed; secondary veins 16–20 on each side of midvein, adaxially shallowly furrowed and impressed or slightly raised. Inflorescence rachis sparsely and shortly hairy. Infructescences 8–18 cm. Cupule wall 1–2 mm thick; bracts spinelike, congested, entirely covering cupule, free or base connate into bundles. Nut 1 per cupule, ovoid, ca. 1.5 cm, pubescent; scar basal, ca. 1 cm in diam. Fl. Apr–Jun, fr. Oct–Dec

of following year.

Broad-leaved evergreen forests on limestone; below 1000 m. W Guangxi, S Guizhou, SE Yunnan [NE Vietnam].

Very similar, if not identical, to Castanopsis indica.

18. Castanopsis longispina (King ex J. D. Hooker) C. C. Huang & Y. T. Chang, Guihaia 12: 1. 1992.

长刺锥 chang ci zhui

Castanopsis tribuloides Smith var. longispina King ex J.

D. Hooker, Fl. Brit. India 5: 623. 1888; C. ferox Spach var. longispina (King ex J. D. Hooker) A. Camus. Trees. Young shoots purple-brown, slightly angulate, tufted puberulent, glabrescent. Petiole 1–1.5 cm; leaf blade oblong to obovate-oblong, 14–24 × 5–8 cm, 1st-year leaf blades papery and abaxially covered with very small stellate cespitose hairs along midvein, base acute to rounded, margin entire, apex shortly acuminate; secondary veins 12–16 on each side of midvein, adaxially ± impressed. Infructescence rachis 4–5 mm thick. Cupule wall ca. 1 mm thick; bracts spinelike, congested, almost completely covering cupule, 1.3–1.5 cm, stiff, free or a few in bundles, puberulent. Nut ellipsoid, 2–3 × ca. 1 cm, glabrescent; scar ca. 8 mm in diam.

Broad-leaved evergreen forests; 800–900 m. SE Xizang (Mêdog Xian) [Bangladesh, NE India, Myanmar, Sikkim].

In India, the species grows at elevations from 200–1200 m.

19. Castanopsis argyrophylla King ex J. D. Hooker, Fl. Brit. India 5: 622. 1888.

银叶锥 yin ye zhui

Trees; branches and leaf blades glabrous. Shoots blackish brown when dry. Petiole 1–2.5 cm; leaf blade elliptic, ovate, lanceolate, or sometimes obovate, 10–20 × 4–7 cm, thickly leathery, abaxially often grayish, adaxially yellow-green when dry, base cuneate to rounded, margin entire, apex mucronate to acuminate; midvein adaxially raised; secondary veins 10–13 on each side of midvein. Infructescences 10–25 cm; rachis covered with pale grayish brown, feltlike, short hairs, glabrescent. Cupule globose, 2.5–3.5 cm in diam., puberulent when young, outside glabrescent, wall 1–1.5 mm thick; bracts spinelike, arranged in discontinuous rings or spirals, 2–6 mm, free or base slightly connate. Nut 1(–3) per cupule, subglobose, 1.5–1.8 cm in diam., densely puberulent. Fl. May–Jun, fr. Oct–Dec of following year.

Broad-leaved evergreen forests; 1000–1500 m. S Yunnan [NE India, Laos, Myanmar, Thailand, Vietnam].

20. Castanopsis tcheponensis Hickel & A.Camus, Notul. Syst. (Paris) 4: 123. 1928.

薄叶锥 bao ye zhui

Trees; branches and leaf blades glabrous. Petiole ca. 1 cm; leaf blade elliptic to ovate-elliptic, 10–15 × 4–6 cm, papery, yellow-green when dry; midvein and secondary veins adaxially slightly raised; secondary veins 9–13 on each side of midvein. Rachis of inflorescences glabrous or very shortly and sparsely mealy puberulent. Female inflorescences 10–25 cm. Cupules shortly stalked when young, subglobose when mature, ca. 3 cm in diam., blackish brown when dry, wall ca. 1 mm thick; bracts spinelike, almost entirely covering cupule, 7–12 mm, slender, free or a few in bundles, glabrous, without scalelike trichomes, basally blackish brown when dry, apically yellowish brown. Nut 1 per cupule, broadly ovoid to subglobose, 1.5–1.8 × 1.2–1.4 cm, glabrous or subglabrous; scar covering more than 3/4 of nut. Fl. Mar–Apr, fr. Oct–Nov.

Broad-leaved evergreen forests; 900–1400 m. S Yunnan [Laos, Myanmar, Vietnam].

21. Castanopsis megaphylla Hu, Bull. Fan Mem. Inst. Biol., Bot. 10: 85. 1940.

大叶锥 da ye zhui

Trees; bud scales, young shoots, and rachis of inflores-cences grayish brown puberulent with small, lamellate, waxy scalelike trichomes. First-year branchlets thick, black-brown when dry, lenticellate; lenticels yellowish brown. Petiole 2–3 cm; leaf blade elliptic, sometimes obovate-elliptic, 26–45 × 8–18 cm, thinly leathery, abaxially midvein and secondary veins stellate puberulent and with small lamellate waxy scalelike tri-chomes when young, surface with a thick tight layer of lamellate waxy scalelike trichomes with age, buff when dry, base cuneate to rounded and often oblique, margin entire, apex obtuse to mucronate; midvein adaxially impressed; secondary veins 16–20 or more on each side of midvein. Female inflorescence ca. 28 cm; rachis densely tawny to dusky puberulent. Infructescence rachis 6–8 mm thick, sparsely lenticellate. Cupule sub-globose, ca. 2.5 cm in diam., outside sparsely puberulent and with waxy scalelike trichomes, basally blackish brown, wall ca. 3 mm thick near base and ca. 1 mm thick near apex; bracts of young cupules spinelike, congested, yellowish brown when dry. Nut subglobose, 1 per cupule, ca. 1 cm in diam., pubescent when young. Fl. May–Jul, fr. of following year.

? Broad-leaved evergreen forests; 1100–1500 m. SE Yunnan (Pingbian Miaozu Zizhixian).

May be the same as the earlier published Vietnamese Castanopsis chevalieri Hickel & A. Camus.

22. Castanopsis nigrescens Chun & C. C. Huang in C. C. Huang & Y. T. Chang, Guihaia 16: 301. 1996.

黑叶锥 hei ye zhui Trees usually 8–15 m tall. First-year branchlets dull blackish brown, often sparsely and thinly grayish pruinose. Petiole 1–2 cm; leaf blade ovate, ovate-elliptic, or rarely lanceolate, 8–15 × 3–6 cm, leathery, abaxially covered with yellowish brown, tightly adnate layers of waxy scalelike trichomes when young, but layers becoming grayish, sparser and thinner with age, adaxially blackish brown to brownish black when dry, base rounded and symmetric or slightly oblique, margin entire, apex acuminate to mucronate; midvein at least from middle to apex adaxially slightly impressed or rarely flat; secondary veins 10–13 on each side of midvein, slender, evident, abaxially slightly raised. Rachis of inflorescences grayish puberulent, glabrescent. Infructescences 5–15 cm; rachis 5–7 mm thick, with cupules crowded from middle to apex. Cupule subglobose, 4–4.5 cm in diam., outside grayish to yellowish gray puberulent, inside brown long tomentose, wall 3–5 mm thick; bracts spinelike, very dense, 1–1.6 cm, grayish to yellowish gray puberulent, free or connate at base or to apical 1/2 and in bundles, usually united to cristate rings. Nut 1 per cupule, broadly ovoid, ca. 2.5 cm, densely pubescent, apex acute; scar covering ca. 1/3 of nut. Fl. May–Jun, fr. Set-Oct of following year.

? Mixed mesophytic and broad-leaved evergreen forests; 200–1000 m. S Fujian, Guangdong, S Guangxi, S Hunan, Jiangxi (Dingnan Xian, Ningdu Xian).

23. Castanopsis damingshanensis S. L. Mo ex C. C. Huang & Y. T. Chang, Guihaia 16: 300. 1996.

大明山锥 da ming shan zhui

Trees 5–9 m tall. Branches glabrous. Petiole 3–6 mm; leaf blade ovate to broadly elliptic, 5–8 × 2.5–4 cm, leathery, concolorous, abaxially sparsely covered with a thin layer of small, yellowish, lamellate scalelike trichomes, base broadly cuneate, margin entire and subconvolute, apex obtuse, rounded, or mucronate; midvein adaxially flat but slightly raised from base to middle; secondary veins 7–10 on each side of midvein. Infructescences few fruited, glabrous; rachis ca. 5 mm thick. Cupule broadly ovoid, 2–2.5 cm in diam., outside and bracts puberulent and with scalelike trichomes, wall 1–2 mm thick; bracts spinelike, basal bracts with a few connected to cristate rings. Nut 1 per cupule, broadly conical, ca. 1.2 cm in diam., brownish pubescent; scar basal, ca. 1 cm in diam. Fr. Oct–Nov.

? Broad-leaved evergreen forests; 1100–1400 m. C Guangxi (Daming Shan).

24. Castanopsis jianfenglingensis Duanmu in W. C. Cheng & al., Sci. Silvae 8: 187. 1963.

尖峰岭锥 jian feng ling zhui

Trees ca. 20 m tall; branchlets and young leaf blades abaxially covered with simple or stellate hairs. Petiole 2.5–3.5 cm, very short when young; leaf blade ovate-elliptic, oblong, or rarely broadly ovate, 12–24 × 5–8

cm, thickly papery, concolorous, base broadly cuneate and often slightly inaequilateral, margin entire or rarely subcrenate from middle to apex, apex caudate; midvein adaxially slightly impressed or rarely flat to slightly raised from base to middle; secondary veins 12–14 on each side of midvein. Infructescences 5–15 cm; rachis 2–3 mm thick. Cupule subglobose, 2–3 cm in diam., wall 1–2 mm thick; bracts spinelike, 3–6 mm, free but usually some in bundles. Nut ellipsoid, 1.2–2 cm, densely rust-colored pubescent; scar basal, 8–10 mm in diam. Fr. Oct–Nov.

? Broad-leaved evergreen forests; 500–800 m. SW Hainan (Dongfang Xian).

25. Castanopsis remotidenticulata Hu, Acta Phytotax. Sin. 1: 104. 1951.

疏齿锥 shu chi zhui

Trees ca. 25 m tall. Branches glabrous; 1st-year branchlets purplish brown. Petiole to 1 cm; leaf blade oblong to obovate-elliptic, 6–12 × 3–4 cm, firmly papery, abaxially covered with tight grayish waxy scalelike trichomes, base broadly cuneate to rounded and often slightly oblique, margin sparsely serrate, apex acuminate to sometimes caudate; midvein and secondary veins adaxially with small ribs; secondary veins 10–13 on each side of midvein. Infructescences

7–10 cm; rachis 4–5 mm thick. Cupule subglobose, 3–3.5 cm in diam., wall 1–2 mm thick; bracts spinelike,

3–6 mm, free but a few in bundles and transversely united to discontinuous cristate rings. Nut 1 per cupule, flat-conical, 1.8–2.4 cm, puberulent; scar basal, 1.8–2.4 cm in diam. Fl. Apr–May, fr. Sep–Nov.

? Broad-leaved evergreen forests; 1000–2200 m. C to SE Yunnan. 26. Castanopsis tonkinensis Seemen, Bot. Jahrb. Syst. 23 (Beibl. 57): 55. 1897.

公孙锥 gong sun zhui

Trees 10–20 m tall. Branches glabrous. Petiole 1–2 cm; leaf blade lanceolate, 6–13 × 1.5–4 cm, membranous, abaxially greenish and sparsely covered with reddish brown, small, lamellate, waxy scalelike trichomes when young, glabrous at maturity, adaxially deep green, base narrowly cuneate, decurrent on petiole, and symmetric or slightly oblique,margin entire, apex caudate to sometimes shortly acute; midvein adaxially somewhat impressed and villous but glabrescent; secondary veins 9–13 on each side of midvein. Female inflorescence ca.

20 cm. Cupule broadly ellipsoid, ovoid, or rarely subglobose, 2–3 cm in diam., dark brownish black when dry, outside glabrescent, wall 0.5–1 mm thick; bracts spinelike, rarely entirely covering cupule, 6–10 mm, glabrescent. Nut 1 per cupule, narrowly conical to broadly ellipsoid, 0.9–1.2 cm in diam., densely brown pilose; scar basal, 8–9 mm in diam. Fl. May–Jun, fr. Sep–Oct of following year. Broad-leaved evergreen forests; below 2000 m. Guangdong, SW Guangxi, Hainan, SE Yunnan [NE Vietnam].

Most Chinese collections are var. tonkinensis, but some specimens from Guangxi and SE Yunnan may be var. laocaiensis Luong, which has broader and thicker leaf blades, and bracts of cupules thicker and shorter than those of var. tonkinensis.

27. Castanopsis amabilis W. C. Cheng & C. S. Chao in W. C. Cheng & al., Sci. Silvae 8: 5. 1963.

南宁锥 nan ning zhui

Castanopsis amabilis var. brevispinosa W. C. Cheng & C. S. Chao.

Trees ca. 20 m tall; branches and rachis of inflorescences glabrous. Petiole 0.5–1.5 cm; leaf blade lanceolate to narrowly ovate, 7–12 × 2–3.5 cm, firmly papery, abaxially with tight layers of yellowish brown scalelike trichomes when young, often grayish with age, base broadly cuneate and symmetric or slightly oblique, margin entire or with 1–3 shallow teeth, apex caudate; midvein adaxially shallowly impressed or occasionally flat; secondary veins 9–13 on each side of midvein, slender, evident. Female inflorescence ca. 26 cm. Infructescence rachis 2–3 mm thick. Cupule broadly ellipsoid to subglobose, 2.2–2.8 cm in diam., outside grayish pubescent or with yellowish brown waxy scalelike trichomes, wall ca. 1 mm thick and exterior visible through bracts; bracts spinelike, 4–8 mm, dispersed or subannular. Nut 1 per cupule, broadly ovoid to subglobose, 0.8–1.2 cm in diam., glabrous; scar basal, 7–10 mm in diam. Fl. Aug–Oct, fr. Oct–Dec of following year.

? Broad-leaved evergreen forests; 300–900 m. SW Guangxi.

28. Castanopsis boisii Hickel & A. Camus, Bull. Soc. Bot. France 68: 396. 1921.

榄壳锥 lan qiao zhui

Castanopsis hamata Duanmu; C. megaphyllya Hu (1948 [1949]), not C. megaphylla Hu (1940).

Trees ca. 25 m tall; branchlets, petiole of young leaves, and mature leaf blades abaxially pubescent and with usually glabrescent yellowish brown to reddish brown, small, lamellate, waxy scalelike trichomes. Petiole 1.5–2 cm; leaf blade ovate-elliptic, ovate, or narrowly oblong, 9–18 × 4–6 cm, thickly papery, base rounded to shortly cuneate and symmetric or slightly oblique, margin entire or rarely from middle to apex shallowly undulate, apex acuminate; midvein adaxially impressed or sometimes slightly raised or flat from base to middle; secondary veins 13–17 on each side of midvein. Infructescence 27 cm; rachis from base to middle 2–3.5 mm thick. Cupule ellipsoid to broadly obovoid, 2.5–3 cm in diam., outside and bracts pubescent and with usually glabrescent yellowish brown to reddish brown small lamellate waxy scalelike trichomes, wall 0.5–1 mm thick and exterior visible through bracts; bracts spinelike, 8–10 mm, greatly variable in thickness, base connate into scattered bundles. Nut 1(or 2) per cupule, broadly ovoid, 1.2–1.4 × 0.9–1.2 cm, puberulent; scar

basal, 8–10 mm in diam. Fl. Jun–Aug, fr. Oct–Nov of following year.

Broad-leaved evergreen forests; 1000–1500 m. Guangdong, SW Guangxi, Hainan, SE Yunnan [NE Vietnam].

29. Castanopsis densispinosa Y. C. Hsu & H. W. Jen, Acta Phytotax. Sin. 13(4): 16. 1975.

密刺锥 mi ci zhui

Trees ca. 15 m tall. Branchlets glabrous, densely lenticellate; lenticels small, grayish. Petiole 1–2 cm; leaf blade ovate to oblong-lanceolate, 13–18 × 3.5–6 cm, green on both surfaces, glabrous, margin coarsely dentate but entire on leaves borne apically on branchlet; midvein abaxially raised and adaxially impressed; secondary veins 10–13 on each side of midvein; tertiary veins abaxially conspicuous. Infructescences to 18 cm. Cupule 2–3 cm in diam., splitting into 3 segments, inside densely grayish villous, wall 1–2 mm thick; bracts spinelike, densely covering outside of cupule, ca.

1 cm, free but basal bracts in bundles. Nut 1 per cupule, ovoid, 1–1.4 × ca. 1 cm, sparsely brownish puberulent, apex obscurely 4- or 5-ridged. Fr. Dec.

? Broad-leaved evergreen forests; ca. 1700 m. Yunnan (Jinping Xian).

30. Castanopsis ledongensis C. C. Huang & Y. T. Chang, Guihaia 16: 301. 1996.

乐东锥 le dong zhui

Trees ca. 18 m tall; young shoots and rachis of male in-florescences pubescent, with reddish brown, small, lamellate, waxy scalelike trichomes, glabrescent. Branchlets dull brown to brownish black. Petiole ca. 1 cm or rarely longer; leaf blade oblong to sometimes obovate-elliptic, 5–9 × 2–3.5 cm, when young abaxially covered with slightly tight layers of reddish brown, small, lamellate scalelike trichomes and with sparsely but soon glabrescent pilose hairs along midvein, base acute and sometimes slightly asymmetric, margin somewhat crenate, apex mucronate to shortly caudate; midvein adaxially impressed; secondary veins 9–12 on each side of midvein. Infructescence ca. 16 cm; rachis 1.5–2 mm thick, glabrous. Cupule subglobose, to ca. 2 cm in diam., splitting into 2 segments, basally abruptly narrowed and slightly decurrent to shortly stalked, wall ca. 1 mm thick; bracts spinelike, 3–5 mm or rarely longer, 3- or 4-angular in cross section, base wide and sometimes several transversely united in a cockscomblike pattern, puberulent and with brownish scalelike trichomes. Nut broadly conical, 1–1.2 cm in diam., apex sparsely puberulent; scar basal, ca. 1 cm in diam. Fr. Oct–Nov.

? Broad-leaved evergreen forests; ca. 800 m. Hainan (Ledong Xian).

31. Castanopsis subuliformis Chun & C. C. Huang in C. C. Huang & Y. T. Chang, Guihaia 16: 301. 1996.

钻刺锥 zuan ci zhui

Trees ca. 25 m tall. First-year branchlets dark brown, glabrous. Petiole ca. 1.5 cm; leaf blade oblong to lanceolate, 7–14 × 3–5 cm, firmly papery, abaxially covered with reddish brown to yellowish brown, small, lamellate waxy scalelike trichomes but soon glabrescent, base rounded to very acute and sometimes slightly inaequilateral, margin from middle to apex with shal-low teeth or rarely entire, apex acute to caudate; midvein adaxially impressed; secondary veins 11–14 on each side of midvein. Infructescences ca. 11 cm; rachis 3–4 mm thick, glabrous. Cupules blackish brown, globose, 1.5–2 cm in diam., wall ca. 1 mm thick; bracts spinelike, scattered, conical, 2–5 cm, yellowish gray pubescent and with small, lamellate, waxy scalelike trichomes, base wide. Nut 1 per cupule, slightly flat-conical, ca. 1.2 × 1.5 cm, densely brown pilose; scar basal, ca. 1.3 cm in diam. Fr. Dec.

? Broad-leaved evergreen forests; 700–900 m. Guangdong, Guangxi.

32. Castanopsis oblonga Y. C. Hsu & H. W. Jen, Acta Phytotax. Sin. 13(4): 19. 1975.

矩叶锥 ju ye zhui

Trees 8–10 m tall. First-year branchlets blackish brown when dry. Petiole 6–10 mm; leaf blade ovate, broadly elliptic, narrowly elliptic, or lanceolate, 6–9 × 2–3.5 cm, thickly papery, brittle when dry, puberulent along midvein and abaxially reddish brown and with a tight layer of scalelike trichomes when young, brownish gray with age, base broadly cuneate to acute and symmetric, margin somewhat crenate to dentate or entire, apex acuminate, caudate, or obtuse; midvein adaxially flat to slightly impressed but usually slightly raised from base to middle; secondary veins 10–14 on each side of midvein. Infructescences 5–10 cm; rachis 2–4 mm thick. Cupule broadly obovoid, ca. 3 cm in diam., dark grayish brown when mature and dry, wall ca. 1 mm thick and exterior ± visible through bracts; bracts spinelike, 4–7 mm, free and evenly scattered or connate at base into short rings, basally gray puberulent, apically yellowish brown and glabrous. Nut broadly conical, 1–1.8 cm in diam., puberulent; scar basal, 0.8–1.4 cm in diam. Fr. Oct–Nov.

? Broad-leaved evergreen forests; ca. 2000 m. SE Yunnan (Yuanjiang Xian).

33. Castanopsis chinensis (Sprengel) Hance, J.Linn. Soc., Bot. 10: 201. 1868.

锥 zhui

Castanea chinensis Sprengel, Syst. Veg. 3: 856. 1826; Castanopsis remotiserrata Hu.

Trees 10–20 m tall; branches and leaf blades glabrous. Petiole 1.5–2 cm; leaf blade lanceolate to rarely ovate, 7–18 × 2–5 cm, thickly papery to nearly leathery, concolorous, base rounded to acute, margin at least from middle to apex serrate, apex caudate; midvein and secondary veins adaxially raised; secondary veins 9–12 on each side of midvein. Female inflorescences borne on apical part of 1st-year branchlets; flowers 1 per cupule. Infructescences 8–15 cm. Cupule globose, 2.5–3.5 cm in diam., usually splitting into 3–5 segments, outside densely grayish brown puberulent when young

but glabrescent, inside densely brown villous, wall 1–1.5 mm thick; bracts spinelike, almost entirely covering outside cupule, 6–12 mm, connate into bundles from base to nearly middle. Nut conical, 1.2–1.6 × 1–1.3 cm, glabrous or rarely puberulent from middle to apex; scar basal, 8–10 mm in diam. Fl. May–Jul, fr. Sep–Nov of following year.

Mixed and broad-leaved evergreen forests; below 1500 m. Guangdong, Guangxi, SW Guizhou, Hunan, SE Yunnan [Vietnam]. 34. Castanopsis eyrei (Champion ex Bentham) Tutcher, J. Linn. Soc., Bot. 37: 68. 1905.

甜槠 tian zhu

Quercus eyrei Champion ex Bentham, Hooker’s J. Bot. Kew Gard. Misc. 6: 114. 1854; Castanopsis asymetrica H. Léveillé; C. brachyacantha Hayata; C. caudata Franchet; C. chingii A. Camus; C. eyrei var. brachyacantha (Hayata) C. F. Shen; C. incana A. Camus; C. neocavaleriei A. Camus; Lithocarpus eyrei (Champion ex Bentham) Rehder; Pasania eyrei (Champion ex Bentham) Oersted; Q. castanopsis H.

Léveillé; Q. cavaleriei H. Léveillé & Vaniot; Q. cepifera H. Léveillé; Q. trinervis H. Léveillé; Shiia brachyacantha (Hayata) Kudo & Masamune; Synaedrys brachyacantha (Hayata) Koidzumi.

Trees 8–20 m tall; branches glabrous. Petiole 0.7–1.5 cm or rarely longer; leaf blade lanceolate, ovate, ovate-elliptic, or oblong, 5–13 × 1.5–5.5 cm, leathery, abaxially reddish brown to dark reddish brown when dry and with brownish to silver-gray, membranous scalelike trichomes, base slightly decurrent on petiole and inaequilateral or sometimes symmetric, margin entire or with few shallow teeth from middle to apex, apex long acuminate; midvein at least from base to middle adaxially slightly raised; secondary veins 7–11 on each side of midvein, very slender, evident. Infructescence rachis 2–5 mm thick, glabrous or glabrescent. Cupule broadly ovoid to subglobose, 2–3 cm in diam., splitting into 2–4 segments, outside and bracts grayish to yellowish gray puberulent, apically acute to obtuse, wall ca. 1 mm thick; bracts spinelike, usually entirely covering outside of cupule, more densely toward apex but if cupule subglobose then basally glabrous and spines sparser, 4–10 mm but apical ones shorter. Nut 1 per cupule, broadly conical, 0.8–1.4 cm in diam., glabrous; scar basal, 8–10 mm in diam. Fl. Apr–Jun, fr. Sep–Nov of following year.

? Mixed and broad-leaved evergreen forests; 300–1700 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Qinghai, Sichuan, Taiwan, Xizang, Zhejiang.

35. Castanopsis ferox (Roxburgh) Spach, Hist. Nat. Vég. 11: 185. 1842.

思茅锥 si mao zhui Quercus ferox Roxburgh, Fl. Ind., ed. 1832, 3: 639. 1832; C. tribuloides (Smith) A. de Candolle var. ferox King ex J. D. Hooker.

Trees 10–20 m tall. Branches glabrous. Petiole 0.8–1.2 cm; leaf blade lanceolate, oblong, or rarely ovate, 8–16 × 2–5 cm, leathery, abaxially with a grayish tight layer of scalelike trichomes, rarely concolorous, base broadly cuneate to cuneate, inaequilateral or symmetric, margin entire, apex acute to acuminate; midvein at least from base to middle adaxially slightly raised or from middle to apex flat to slightly impressed; secondary veins 9–14 on each side of midvein. Rachis of inflorescences tomentulose. Female inflorescence ca. 20 cm. Perianth puberulent. Infructescence rachis glabrous. Cupule globose to rarely broadly ovoid, 2–2.8 cm in diam., outside and bracts brown to grayish brown pubescent and with waxy scalelike trichomes, wall to 1 mm thick; bracts spinelike, 4–8 mm, base connate into bundles or united to discontinuous rings. Nut subglobose, 0.8–1.2 × 0.9–1.2 cm, glabrous, apex pointed; scar basal, 8–10 mm in diam. Fl. Aug–Oct, fr. Sep–Nov of following year.

Broad-leaved evergreen forests; 700–2000 m. SE Xizang, S to SW Yunnan [Bangladesh, NE India, Laos, Myanmar, Sikkim, N Thailand, Vietnam].

36. Castanopsis fargesii Franchet, J. Bot. (Morot) 13: 195. 1899.

栲 kao

Castanopsis argyracantha A. Camus; C. cryptoneuron (H. Léveillé) A. Camus ex Rehder; C. taiwaniana Hayata; Pasania ischnostachya Hu; Quercus cryptoneuron H. Léveillé; Q. pinfaensis H. Léveillé & Vaniot.

Trees 10–30 m tall; bud scales, young branchlets from middle to apex, petiole of young leaf blades, and leaf blades abaxially covered with glabrescent, rust-colored, small, lamellate, waxy scalelike trichomes. Branches glabrous. Petiole 1–2 cm; leaf blade oblong, lanceolate, or rarely ovate, 7–15 × 2–5 cm, abaxially covered with a thick and mealy layer of scalelike trichomes, reddish brown to yellowish brown when young, but tawny with age, base rounded to broadly cuneate and sometimes inaequilateral, margin entire or sometimes with few shallow teeth from middle to apex, apex acute to acuminate; midvein at least from middle to apex adaxially impressed; secondary veins 11–15 on each side of midvein. Female inflorescences solitary, glabrous, to 30 cm; cupules scattered on rachis. Infructescence rachis 1.5–3 mm thick. Cupule globose to broadly ovoid, 2.5–3 cm in diam., splitting irregularly, outside and bracts whitish gray to brownish puberulent or with reddish brown waxy scalelike trichomes and sparse pubescence, wall ca. 1 mm thick; bracts spinelike, 8–10 mm, connate and in bundles

basally or rarely from base to middle. Nut 1 per cupule, conical and 1–1.5 × 0.8–1.2 cm to subglobose and 0.8–1.4 cm in diam., glabrous; scar basal, 8–10 mm in diam. Fl. Apr–Jun and Aug–Oct, fr. Apr–Oct of following year.

? Broad-leaved evergreen forests; 200–2100 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Si-chuan, Taiwan, Yunnan, Zhejiang.

37. Castanopsis ouonbiensis Hickel & A. Camus, Bull. Soc. Bot. France 68: 398. 1921.

屏边锥 ping bian zhui

Trees ca. 25 m tall; 1st-year branchlets, petioles, and rachis of infructescences yellowish brown to dark dirty brown tomentulose. Petiole 2–2.5 cm; leaf blade elliptic to obovate-elliptic, 10–18 × 4–6 cm, subleathery, abaxially puberulent and with early glabrescent, reddish brown, mealy, waxy scalelike trichomes, base broadly cuneate to rounded and often inaequilateral, margin entire, apex shortly acuminate to acute; midvein adaxially impressed; secondary veins 10–15 on each side of midvein. Female inflorescences 10–20 cm. Infructescences 10–16 cm; rachis slender. Cupule subglobose to broadly ellipsoid, 2.5–3 cm in diam., outside and bracts rusty brow puberulent and with lamellate, waxy scalelike trichomes, wall 1–2 mm thick and exterior visible through bracts; bracts spinelike, 4–8 mm, in separated bundles, usually connate at middle or apically, rarely connate basally. Nut conical, 1.4–1.8 × 1.2–1.4 cm, glabrous; scar basal, 1–1.2 cm in diam. Fl. Oct–Nov, fr. Oct–Nov of following year.

Broad-leaved evergreen forests; 1100–1600 m. SE Yunnan [N Vietnam].

38. Castanopsis jucunda Hance, J. Bot. 22: 230. 1884.

秀丽锥 xiu li zhui

Castanopsis formosana (Skan) Hayata; C. tribuloides (Smith) A. de Candolle var. formosana Skan.

Trees 10–26 m tall; 1st-year branchlets and leaf blades adaxially brownish black when dry; bud scales, branchlets, petiole of young leaf, mature leaf blades abaxially, and rachis of inflorescences sparsely covered with glabrescent, reddish brown, waxy scalelike trichomes. Petiole 1–2.5 cm; leaf blade ovate, ovate-elliptic, or oblong, sometimes obovate to obovate-elliptic, (4–)10–18 × (2–)4–8 cm, base rounded to broadly cuneate and inaequilateral or symmetric, margin at least from middle to apex serrate or rarely undulate, apex acute to acuminate; midvein adaxially impressed; secondary veins (5–)8–11 on each side of midvein. Female inflorescences axillary, solitary. Infructescences 10–15 cm. Cupule subglobose, 2.5–3.5 cm in diam., splitting into 3–5 segments, outside and bracts pubescent and with grayish brown, lamellate, waxy scalelike trichomes especially when young, wall 0.5–1(–1.5) mm thick; bracts spinelike, 6–10 mm, free but a few in bundles, sometimes transversely united to discontinuous rings. Nut broadly conical to ovoid, 1–2 × 1–1.5 cm, glabrous or glabrescent; scar basal, 1–1.2(–1.5) cm in diam. Fl. Apr–May, fr. Aug–Oct of following year.

Broad-leaved evergreen forests; below 1500 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, E to SC Taiwan, Yunnan, Zhejiang [Vietnam].

39. Castanopsis hupehensis C. S. Chao in W. C. Cheng & al, Sci. Silvae 8(2): 187. 1963.

湖北锥 hu bei zhui

Trees 10–20 m tall; 1st-year branchlets and bud scales glabrous. Petiole 3–7(–10) mm; leaf blade lanceolate to oblong, sometimes obovate-elliptic to oblanceolate, 6–11 × 1.5–3.5 cm, subleathery, abaxially brownish when young and dry but grayish with age and with tight layers of scalelike trichomes, base broadly cuneate and often slightly oblique, margin serrate from middle to apex or entire, apex acuminate to abruptly narrowly caudate; midvein adaxially slightly impressed; secondary veins 10–13 on each side of midvein. Rachis of inflorescences glabrous, without waxy scalelike trichomes. Infructescence rachis 1.5–3 mm thick. Cupule ellipsoid to subglobose when mature, 2–2.2 cm in diam., outside grayish to yellowish brown puberulent, basally often shortly stalked, wall ca. 1 mm thick; bracts spinelike, 4–6 mm, few of them connate into bundles, others transversely united to 4 or 5 cristate rings. Nut 1 per cupule, broadly conical, glabrescent; scar basal, 9–12 mm in diam. Fl. Jun–Sep, fr. Jun–Nov of following year.

? Broad-leaved evergreen forests; 600–1000 m. NE Guizhou, W Hubei, NW Hunan, E Sichuan.

40. Castanopsis fleuryi Hickel & A. Camus, Bull. Soc. Bot. France 68: 395. 1921.

小果锥 xiao guo zhui

Castanopsis microcarpa Hu.

Trees to 10 m tall. Branches glabrous. Petiole 0.8–1.5 cm; leaf blade elliptic, lanceolate, or ovate, 9–20 × 3–7 cm, subleathery, abaxially with tight layers of trichomes, 1st-year leaf blades concolorous, base cuneate to rounded and sometimes oblique, margin entire, apex acuminate; midvein adaxially raised from base to middle and slightly impressed from middle to apex; secondary veins 7–10 on each side of midvein. Infructescences 8–15 cm; rachis 2–4 mm thick, yellowish gray pubescent. Cupule ellipsoid to broadly ovoid, 7–11 mm in diam., outside sparsely covered with spinelike bracts and yellowish gray to grayish brown pubescence and waxy scalelike trichomes, basally shortly stalked, apically narrowly pointed, wall to 0.5 mm thick; bracts 1–3 mm, in rows, 3- or 4-angular in cross section, free or a few connected at base to continuous or discontinuous rings. Nut 1 per cupule, broadly conical, 8–12 × 7–10 mm, glabrescent when

mature; scar basal, 6–8 mm in diam. Fl. May–Jul or

Oct–Nov, fr. Oct–Nov of following year.

Broad-leaved evergreen forests; 600–2400 m. S to SW Yunnan [Laos, Vietnam].

41.Castanopsis rufotomentosa Hu,Bull. Fan Mem. Inst. Biol., n.s. 1: 223. 1948 [1949].

红壳锥 hong qiao zhui

Trees 18–25 m tall; 1st-year branchlets, petioles, and leaf blades abaxially covered with early glabrescent, dark reddish brown, small, lamellate, waxy scalelike trichomes. Petiole 2–8 mm; leaf blade narrowly lanceolate to oblong, 10–20 cm × 2–4 mm, leathery, base acute to rounded, margin entire or rarely with a few shallow teeth from middle to apex, apex narrow and obtuse; midvein and secondary veins adaxially impressed; secondary veins 14–18 on each side of midvein. Infructescences 8–15 cm; rachis covered with mealy, waxy scalelike trichomes. Cupule globose, 1.2–2.2 cm in diam., outside and bracts covered with early glabrescent, dark reddish brown, smal,l lamellate, waxy scalelike trichomes, wall ca. 1 mm thick and exterior visible through bracts; bracts spinelike, 3–6 mm, free or several with base connate into bundles. Nut 1 per cupule, oblate-globose to subglobose, ca. 1 cm in diam., glabrous, apex pointed; scar basal, ca. 9 mm in diam. Fr. Nov–Dec.

? Broad-leaved evergreen forests; ca. 1300 m. SE Yunnan (Xichou Xian).

42. Castanopsis tribuloides (Smith) A. de Candolle in Hance, J. Bot. 1: 182. 1863.

蒺藜锥 ji li zhui

Quercus tribuloides Smith in Rees, Cycl. 29: Quercus no. 13. 1814; Castanea tribuloides (Smith) Lindley. Trees 5–10 m tall; young branchlets and young leaf blades abaxially pubescent and with glabrescent, rusty brown, waxy scalelike trichomes. Petiole 1–1.5 cm; leaf blade elliptic to ovate, 9–16 × 3.5–5 cm, abaxially reddish brown but may become gray to grayish brown with age, base acute to rounded, margin entire or rarely with 1 or 2 teeth, apex acute; midvein adaxially impressed; secondary veins 11–14 on each side of midvein, sometimes impressed. Infructescences ca. 25 cm; rachis slender. Cupules loosely arrange, globose to ellipsoid, 1.6–2.2 cm in diam., outside covered with brownish, small, lamellate, waxy scalelike trichomes, sometimes pubescent, wall to 1 mm thick; bracts spinelike, sparsely covering cupule, 3–5 mm, slender, free but a few in bundles. Nut 1 per cupule, broadly conical, 1.5–2 × 1–1.6 cm, glabrous; scar basal, 8–10 mm in diam. Fl. Apr–May, fr. Sep–Oct of following year.

Broad-leaved evergreen forests; ca. 1300 m. SE Xizang (Mêdog Xian), SW Yunnan [N India, Myanmar, Nepal, N Thailand].

43. Castanopsis wenchangensis G. A. Fu & C. C. Huang, Acta Phytotax. Sin. 27: 151. 1989. 文昌锥 wen chang zhui

Trees 5–8 m tall; branches, bud scales, and rachis of inflorescences glabrous. Petiole 1–2 cm; leaf blade lanceolate to ovate, (3–)5–9(–12) × (1.2–)2–3.5(–6) cm, leathery, abaxially with thin greenish layers of adnate scalelike trichomes when young, concolorous with age, base rounded to acute, margin with shallow to deep teeth, apex acuminate to rarely acute; midvein at least from base to middle adaxially raised to rarely slightly impressed; secondary veins 6–10 on each side of midvein, adaxially raised. Female inflorescences 3–8 cm. Infructescences 4–5 cm, with 1–6 fruit; rachis 1–1.5 mm thick. Cupule subglobose, 1.5–2 cm in diam., completely enclosing nut, irregularly splitting into 3 or 4 segments, outside sparsely pubescent and with small, lamellate, waxy scalelike trichomes, basally without bracts, wall 1–2 mm thick; bracts spinelike, scattered, free or a few in bundles, 2–4 mm. Nut subglobose, 1.3–1.4 cm in diam., puberulent; scar basal, 1–1.2 cm in diam. Fl. Jul–Aug, fr. Oct–Dec of following year.

? Broad-leaved evergreen forests. Hainan (Wenchang Xian).

44. Castanopsis delavayi Franchet, J. Bot. (Morot) 13: 194. 1899.

高山锥 gao shan zhui

Castanopsis tsaii Hu; Synaedrys delavayi (Franchet) Koidzumi.

Trees ca. 20 m tall. Branches glabrous. Petiole 0.7–1.5 cm; leaf blade obovate, obovate-elliptic, ovate, or elliptic, 5–13 × 3–7 cm, subleathery, abaxially sparsely covered with yellowish brown membranous trichomes when young, grayish to silver-gray with age, base acute to rounded, margin serrate or rarely undulate, apex acute to rounded; midvein adaxially raised; secondary veins 6–9 on each side of midvein, slightly raised. Female inflorescence rachis glabrous. Infructescences 10–15 cm; rachis 2–3 mm thick. Cupule ellipsoid when young but broadly ovoid to subglobose with age, 1.5–2 cm in diam. or slightly larger, splitting into 2 or 3 segments, outside with yellowish brown, waxy scalelike trichomes and adnate pubescence, basally long stalked, wall 0.5–1 mm thick; bracts spinelike, 3–6 mm, free or connate at base and transversely united to 3–5 circular or spiral rings, covered with yellowish brown waxy scalelike trichomes and adnate pubescence. Nut broadly ovoid, 1.3–1.4 cm in diam.; scar basal, 6–8 mm in diam. Fl. Apr–May, fr. Sep–Nov of following year.

? Mixed and broad-leaved evergreen forests; 1500–2800 m. Guangdong, SW Guizhou, SW Sichuan, Yunnan.

45. Castanopsis echinocarpa J. D. Hooker & Thomson ex Miquel, Ann. Mus. Bot. Lugduno-Batavi 1: 119. 1863.

短刺锥 duan ci zhui

Castanopsis echinocarpa var. seminuda W. C. Cheng & C. S. Chao; C. longispicata Hu; C. tribuloides (Smith)

A. de Candolle var. echinocarpa (J. D. Hooker & Thomson ex Miquel) King ex J. D. Hooker.

Trees 7–15(–25) m tall. Branches glabrous; branchlets dark brown to brownish black when dry. Petiole 1–1.5 cm; leaf blade elliptic, ovate, or lanceolate, (5.5–)7–12.5(–14.5) × (1.5–)2.5–5(–6) cm, thickly papery, abaxially covered with a very tight layer of scalelike trichomes and reddish brown to yellowish brown when young but grayish brown with age, base rounded to sometimes acute, margin serrate to rarely subentire, apex acuminate to abruptly narrowly caudate; midvein adaxially slightly impressed to rarely nearly flat; secondary veins 9–13 on each side of midvein. Rachis of inflorescences glabrous or glabrescent. Infructescence rachis base 2–3.5 mm thick. Cupule globose, 1.5–2 cm in diam., rarely splitting, wall ca. 1 mm thick; bracts spinelike, thick, 1–3 mm, sometimes no more than tubercles, 3- or 4-angular in cross section, pale grayish brown pubescent, sometimes base loosely connected to discontinuous rings. Nut 1 per cupule, subglobose to conical, 1–1.3 cm in diam., glabrous, apex acute; scar basal, 8–9 mm in diam. Fl. Apr–May, fr. Sep–Oct of following year.

Broad-leaved evergreen forests; 500–2300 m. SE Xizang, S Yunnan [Bangladesh, Bhutan, NE India, Myanmar, Nepal, Thailand, Vietnam].

46. Castanopsis carlesii (Hemsley) Hayata, Icon. Pl. Formo-san. 6 Suppl.: 72. 1917.

米槠 mi zhu

Trees ca. 20 m tall; young shoots and rachis of inflores-cences sparsely covered with reddish brown, lamellate, waxy scalelike trichomes. Petiole to ca. 1 cm, base becoming thick, pillow-shaped; leaf blade lanceolate to ovate, 4–12 × 1–4.5 cm, leathery, abaxially with layers of reddish brown to yellowish brown, slightly adnate, small, lamellate scalelike trichomes when young but grayish brown to silvery with age, base slightly cuneate to broadly so and oblique, margin entire or with a few shallow teeth, apex acuminate to narrowly caudate; midvein adaxially flat to slightly impressed but impressed when dry; secondary veins 8–13 on each side of midvein. Rachis of male catkins glabrous or glabrescent. Infructescence rachis 2–3 mm thick, glabrous. Cupule subglobose to ovoid, 1–1.5 cm, outside yellowish brown to reddish brown feltlike pubescent and covered with waxy scalelike trichomes, wall 0.5(–1) mm thick; bracts spinelike or reduced to tubercles. Nut subglobose to broadly conical, apex shortly pointed; scar basal, ca. 5 mm in diam. Fl. Mar–Jun, fr. Sep–Nov of following year.

? Mixed and broad-leaved evergreen forests, mixed mesophytic forests; below 1700 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang. The variation in cupule spines length is continuous between the two extremes, and it is not always possible to distinguish the two varieties. 1a. Cupules tuberculate or cupules from middle to apex with free scattered spinelike bracts

1–2

mm and from base to middle without bracts

or with very short bracts .............. 46a. var. carlesii 1b. Cupules covered with adnate to fascicled

spinelike bracts 2–5(–7) mm .... 46b. var. spinulosa 46a. Castanopsis carlesii var. carlesii

米槠(原变种) mi zhu (yuan bian zhong)

Quercus carlesii Hemsley, Hooker’s Icon. Pl. 26: t. 2591. 1899; Castanopsis carlesii var. sessilis Nakai; C. cuspidata (Thunberg) Schottky var. carlesii (Hemsley) T. Yamazaki; C. cuspitata var. longicaudata (Hayata) S. S. Ying; C. longicaudata (Hayata) Nakai; C. stipitata (Hayata ex Koidzumi) Nakai; Lithocarpus stipitatus Hayata ex Koidzumi; Q. longicaudata Hayata; Shiia carlesii (Hemsley) Kudo; Synaedrys carlesii (Hemsley) Koidzumi.

Cupules with tubercles or sometimes apically with spinelike bracts ca. 1–2 mm, sometimes adnate but not connate into fascicles.

? Mixed and broad-leaved evergreen forests. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang.

46b. Castanopsis carlesii var. spinulosa W. C. Cheng & C. S. Chao in W. C. Cheng & al., Sci. Silvae 8: 6. 1963.

短刺米槠 duan ci mi zhu

Cupule bracts spinelike, more dense apically, 2–5(–7) mm, longer from middle to apex of cupule, basal bracts connate into bundles.

? Mixed mesophytic forests; 1000–1700 m. Guangxi, Guizhou, Hunan, Sichuan, Yunnan.

47. Castanopsis xichouensis C. C. Huang & Y. T. Chang, Guihaia 10: 2. 1990.

西畴锥 xi chou zhui

Trees ca. 15 m tall. Branches glabrous. Petiole 1.5–2.5 cm; leaf blade elliptic, ovate-elliptic, or rarely obovate-elliptic, 10–15 × 3.5–6 cm, firmly leathery, abaxially reddish brown, sparsely covered with adnate, black-brown, rounded, lamellate, waxy scalelike trichomes when young, grayish and waxy with age, base shortly attenuate to broadly cuneate and sometimes oblique, margin serrate from middle to apex, apex acute; mid-vein adaxially slightly raised; secondary veins 10–13 on each side of midvein. Infructescences dense, 10–30 cm; rachis 6–10 mm thick. Cupule 4–4.5 cm in diam., wall 1–2 mm thick; bracts spinelike, entirely covering outside of cupule, connate at base or to middle 1/2 into 3–7-spined bundles. Nuts (2 or)3 per cupule, broadly conical, ca. 1.2 cm in diam., densely brown puberulent; scar 6–10 mm in diam. Fr. Oct.

? Broad-leaved evergreen forests; 1400–1700 m. Yunnan (Xichou Xian).

48. Castanopsis rockii A. Camus, Bull. Bimenstr. Soc. Linn. Lyon 8: 88. 1929.

龙陵锥 long ling zhui

Castanopsis lunglingensis Hu.

Trees 20–27 m tall. Branches glabrous; young shoots dark purplish red, dark brown when dry. Petiole 1.5–2 cm; leaf blade oblong, narrowly elliptic, or sometimes oblanceolate, 15–25 × 4–7 cm, papery, abaxially with a tight and thin layer of scalelike trichomes, base cuneate and oblique, margin entire, apex acute to acuminate; midvein and secondary veins adaxially slightly raised; secondary veins 14–17 on each side of midvein. Male inflorescences ca. 20 cm; rachis sparsely puberulent. Female flowers 3 per cupule. Infructescence rachis glabrous, 5–8 mm thick, fruit loosely arranged. Cupule subglobose to broadly elliptic, 4–5 cm in diam., outside glabrescent, dark brown when dry, wall 2–3 mm thick; bracts spinelike, entirely covering cupule, 1–1.5 cm, base connate into bundles. Nuts 2 or 3 per cupule, broadly conical, ca. 1.2 mm in diam., densely hairy; scar covering ca. 1/3 of nut. Fr. Oct–Nov.

Mixed and broad-leaved evergreen forests; below 2100 m. S and W Yunnan [Thailand, Vietnam].

49. Castanopsis tessellata Hickel & A. Camus, Bull. Soc. Bot. France 68: 399. 1921.

棕毛锥 zong mao zhui

Trees 10–15 m tall; bud scales, young shoots, petioles, leaf blades abaxially, and rachis of inflorescences densely brown pilose. Petiole 1.5–3 cm; leaf blade oblong, sometimes lanceolate, 15–30 × 5–8 cm, base acute to broadly cuneate and symmetric to sometimes oblique, margin entire, apex long acuminate; midvein adaxially slightly raised and puberulent; secondary veins 16–22 on each side of midvein, slightly impressed. Male flowers spirally arranged on rachis. Female in-florescences crowded from middle to apex of branchlets; flowers 3 per cupule, often with staminodes. Infructescences 10–30 cm. Cupule subglobose, 5–6 cm in diam., base narrowing into a stalk 3–5 mm, wall 1–1.5 mm thick, outside except basally densely covered with spinelike bracts, inside densely yellowish brown pilose, spinelike bracts basally in bundles, yellowish brown pilose. Nuts 2 or 3 per cupule, broadly conical, 1.5–1.8 × 1.5–2 cm, pilose; scar basal, 1–1.5 cm in diam. Fl. Apr–Jun, fr. Sep–Oct.

Broad-leaved evergreen forests; below 500 m. SE Yunnan [C to N Vietnam].

50. Castanopsis kweichowensis Hu, Bull. Fan Mem. Inst. Biol., n.s. 1: 221. 1948 [1949].

贵州锥 gui zhou zhui

Trees ca. 20 m tall; young shoots, petioles, leaf blades abaxially, and rachis of inflorescences densely yellowish brown pilose. Petiole 0.5–1.5 cm, thick; leaf blade narrowly oblong to oblanceolate-oblong, 16–30 × 5–9 cm, firmly papery, base rounded to broadly cuneate and symmetric or oblique, margin apically serrate, apex abruptly acute; midvein and secondary veins adaxially impressed; secondary veins 16–25 on each side of midvein. Female inflorescences 10–25 cm; flowers 3 per cupule. Infructescence rachis 7–12 mm thick. Cupule irregularly globose, 3–3.5 cm in diam., outside pubescent, basally narrowed, wall 2–2.5 mm thick; bracts spinelike, basal bracts often transversely connected to ring but apically in bundles, 5–8 mm, pubescent. Nuts 2 or 3 per cupule, broadly conical, 1.6–2 × 1.6–2.2 cm, tomentose; scar basal, 1–1.5 cm in diam. Fl. May–Jun, fr. Oct–Dec of following year.

? Broad-leaved evergreen forests; 400–800 m. N Guangxi, S Guizhou. Very similar to Castanopsis ceratacantha but with denser cupule spines and larger leaves; it could be conspecific.

51. Castanopsis ceratacantha Rehder & E. H. Wilson in Sargent, Pl. Wilson. 3: 199. 1916.

瓦山锥 wa shan zhui

Castanopsis chuniana W. P. Fang.

Trees 8–15 m tall; 1st-year branchlets and rachis of inflorescences yellowish brown to brownish villous. Petiole ca. 1 cm; leaf blade lanceolate, oblong, or sometimes oblanceolate, 10–18 × 2–5 cm or rarely shorter or wider, papery, 1st-year leaf blades at least along veins abaxially villous and with layers of reddish brown to yellowish brown scalelike trichomes, base broadly cuneate to acute and oblique, margin apically with 2–5 shallow teeth or entire, apex long acuminate to mucronate; midvein adaxially impressed; secondary veins 13–17 on each side of midvein, slightly impressed to raised. Female inflorescences in clusters on apical part of branchlet of previous year; flowers 2 or 3 per cupule. Cupule subglobose, ca. 3 cm in diam., outside and bracts brown villous and with small, lamellate, waxy scalelike trichomes, wall 1–1.5 mm thick; bracts spinelike, to 5 mm, several connate into bundles from middle or apical part, sometimes cristate. Nuts 1 or 2(or 3) per cupule, broadly conical, 1.5–1.8 × 1.4–1.8 cm, pubescent; scar basal, ca. 1 cm in diam. Fl. Apr–May, fr. autumn to early winter of following year.

Mixed and broad-leaved evergreen forests; 1500–2500 m. Guizhou, Hubei, SW Sichuan, Yunnan [Laos, NE Thailand, Vietnam].

52. Castanopsis fabri Hance, J. Bot. 22: 230. 1884.

罗浮锥 luo fu zhui

Castanopsis brevispina Hayata; C. brevistella Hayata & Kanehira ex A. Camus; C. ceratacantha Rehder & E. H. Wilson var. semiserrata (Hickel & A. Camus) A. Camus; C. hickelii A. Camus; C. kusanoi Hayata; C. matsudai Hayata ex A. Camus; C. ninbienensis Hickel & A. Camus; C. quangtriensis Hickel & A. Camus; C. semiserrata Hickel & A. Camus; C. sinsuiensis Kanehira; C. stellatospina Hayata; C. tenuispinula Hickel & A. Camus; C. traninhensis Hickel & A. Camus.

中华人民共和国人力资源和社会保障部令第13号

中华人民共和国人力资源和社会保障部令第 13号 《实施〈中华人民共和国社会保险法〉若干规定》已经人力资源和社会保障部第67次部务会审议通过,现予公布,自2011年7月1日起施行。 部长尹蔚民 二O一一年六月二十九日 实施《中华人民共和国社会保险法》若干规定 为了实施《中华人民共和国社会保险法》(以下简称社会保险法),制定本规定。

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第二条县级以上的地方各级人民代表大会设立常务委员会。 第二章地方各级人民代表大会 第三条地方各级人民代表大会都是地方国家权力机关。 第四条省、自治区、直辖市、自治州、设区的市的人民代表大会代表由下一级的人民代表大会选举;县、自治县、不设区的市、市辖区、乡、民族乡、镇的人民代表大会代表由选民直接选举。 地方各级人民代表大会代表名额和代表产生办法由选举法规定。各行政区域内的少数民族应当有适当的代表名额。 第五条省、自治区、直辖市、自治州、设区的市的人民代表大会每届任期五年。县、自治县、不设区的市、市辖区、乡、民族乡、镇的人民代表大会每届任期三年。 第六条省、自治区、直辖市的人民代表大会根据本行政区域的具体情况和实际需要,在和国家宪法、法律、政策、法令、政令不抵触的前提下,可以制订和颁布地方性法规,并报全国人民代表大会常务委员会和国务院备案。 第七条县级以上的地方各级人民代表大会行使下列职权: (一)在本行政区域内,保证宪法、法律、政策、法令、政令和上级人民代表大会决议的遵守和执行,保证国家计划和国家预算的执行; (二)审查和批准本行政区域的国民经济计划和预算、决算; (三)讨论、决定本行政区域的政治、经济、文化、教育、卫生、民政、民族工作的重大事项; (四)选举本级人民代表大会常务委员会的组成人员; (五)决定省长、副省长,自治区主席、副主席,市长、副市长,州长、副州长,县长、副县长,区长、副区长的人选;

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第三条给予事业单位工作人员处分,应当坚持公正、公平和教育与惩处相结合的原则。 给予事业单位工作人员处分,应当与其违法违纪行为的性质、情节、危害程度相适应。 给予事业单位工作人员处分,应当事实清楚、证据确凿、定性准确、处理恰当、程序合法、手续完备。 第四条事业单位工作人员涉嫌犯罪的,应当移送司法机关依法追究刑事责任。 第二章处分的种类和适用 第五条处分的种类为: (一)警告; (二)记过; (三)降低岗位等级或者撤职; (四)开除。 其中,撤职处分适用于行政机关任命的事业单位工作人员。 第六条受处分的期间为: (一)警告,6个月; (二)记过,12个月; (三)降低岗位等级或者撤职,24个月。 第七条事业单位工作人员受到警告处分的,在受处分期间,不得聘用到高于现聘岗位等级的岗位;在作出处分决定的当年,年度考核不能确定为优秀等次。 事业单位工作人员受到记过处分的,在受处分期间,不得聘用到高于现聘岗位等级的岗位,年度考核不得确定为合格及以上等次。

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向着中国梦,前进! 绥芬河市北寒小学四年级李鸣 梦,通常指人们的理想。每个人都会做梦,每个人都会有自己的梦想。美羊羊的梦想是能够在青青草原上尽情地玩耍,光头强的梦想是把大森林里的树木都砍光。我的梦想,可以是一个漂亮的发卡,一盒可口的点心,是考试后的100分,妈妈对我夸奖的笑容……可今天,老师说,我们每个人不但要有自己的梦,还要有中国梦。 那么,什么是中国梦呢?中国梦是建立在祖国发展基础上的,是对祖国有利的梦,每个人的中国梦都不相同,有大有小,但它们又都有相同之处,那就是都怀着一颗为祖国作出贡献的心,对人民有功利的心。我想中国梦还应该是国家富强,小朋友们都有新衣服穿,有变形金刚、芭比娃娃;民族团结,不会有战争与饥饿;大人们高高兴兴地上班,孩子们能够一块儿在草地上做游戏。但我觉得,最重要的是要建设一个美丽、洁净的家园,那就是天蓝、地绿、水清、气爽的美丽中国。没有地沟油,没有沙尘暴,没有禽流感…… 建设美丽中国,就要从小事做起。在学校,看到地上有纸屑,我会主动弯腰捡起;看到水龙头滴水,我要主动上前把它关紧;看到有人说脏话,我会及时上前制止。春天来了,我拉着爸爸妈妈买树苗,到附近山坡上植树,因为我知道,绿化造林是防治大气污染的一种有效方法。我还要宣传环保知识,动员更多的人加入环保队伍,让天变得更蓝,水变得更绿。我还会在村里钉上一块“爱护环境、人人有责”的警示牌,让全村的人跟我一起行动起来,保护我们的环境、净化我们的环境。我知道我一个人,一家人的力量很微薄,但是如果全中国的孩子都能行动起来,每个中国家庭都行动起来,那就会汇成强大的环保能量,才能建设好美丽中国。 每个中国人都有着自己的中国梦,只要每个人能为这伟大的梦想贡献出自己的力量,哪怕是微不足道,但当所有的贡献汇集到一起,将会是一股多么强大的力量,这股力量会把中国推向更美好的明天,只要人人行动起来,我相信建设美丽祖国这个中国梦将会很快实现。

中国梦-我的梦演讲稿

中国梦我的梦 有一种东西,它承载着人们的希望,虽然他看不见,摸不着,却能在人们心中产生巨大的力量。它,就叫梦想。上帝没有赐予我们翅膀,却赐予了我们会飞的心灵和能够梦想的大脑,使我们有了一双“隐形的翅膀”,带着我们在人生辽阔的天空里自由地翱翔。 今年三月,全国人大和全国政协会议在北京召开,关于“中国梦”的话题也进行过热烈的讨论。每个人或多或少也在思考“中国梦”的问题。 那么,中国梦是什么呢?习近平总书记说“每个人都有理想和追求,都有自己的梦想。实现中华民族的伟大复兴,就是中华民族近代以来最伟大的梦想。这个梦想,凝聚了几代中国人的夙愿,体现了中华民族和中国人民的整体利益,是每一个中华儿女的共同期盼。”是的,每个人都有自己的梦,梦想能够照亮生活,亦能成就未来。而有一个梦,它既是你的梦,也是我的梦,它是万千中华儿女共同的心愿,那就是实现中华民族伟大复兴的中国梦。 每个人的梦都是中国梦的组成部分,我的梦亦如此。都说学生的主业是学习,的确如此,我的梦来自学习。我喜欢在午休时一个人独自读书的静谧;喜欢在难题前不停运用各种公式演算的执着;也喜欢与同学一起为一道题目争执不休的吵吵闹闹。这样所诠释的梦是理想之梦。 梦也来自家庭,对于我的成长,它最有发言权。平日里与父母相处的时间虽然不多,但是一起吃饭时,父亲在饭桌上的高谈阔论;

坐在同一书桌前各自忙碌时,偶尔抬头与母亲随意交谈的闲话,都在影响着我对生活的态度。 有了梦,便要去追寻。三年前,在懵懂的状态下,在家长的引导下选好了脚下的路—上高中、考大学。渐渐地,考大学也真正成为了我的梦。清晰地记得,老师在中考前说“我们将做出人生第一次重大选择”,面对众多的学校,我固执地选择了朝中,心中似乎在坚守着什么。后来,随着一个个目标的制定,无数次为“自己的事”的奋斗,我逐渐明白,我是在坚守着自己的梦,并且开始懂得为了理想和追求而努力拼搏。其实走到现在,蓦然回首,才发现人生是一个条件从句,梦的方向并无过多区别,而我们最终选择的那个梦,正是因为被自己选择,被自己填充,才显得与众不同,才显得适合自己。 我相信,只要坚持不懈,努力拼搏,梦想就一定能够成真。我的梦,我们的梦,铸就了实现国富民强的中国梦。可以想想,如果中国民族实现了伟大复兴,那么世界梦一定会更加流光溢彩。

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中华人民共和国劳动和社会保障部令 第2号 依据《社会保险费征缴暂行条例》(中华人民共和国国务院令第259号),劳动和社会保障部制定了《社会保险费申报缴纳管理暂行办法》,现予发布施行。 劳动和社会保障部部长 张左己 一九九九年三月十九日 社会保险费申报缴纳管理暂行办法 第一条 为规范社会保险费的申报和缴纳管理工作,根据《社会保险费征缴暂行条例》(以下简称条例)的规定,制定本办法。 第二条 本办法规定的社会保险费是指基本养老保险费、失业保险费和基本医疗保险费。工伤保险费和生育保险费的申报、缴纳可参照本办法执行。 第三条 缴费单位进行缴费申报和社会保险经办机构征收社会保险费适用本办法。由税务机关征收社会保险费的缴纳管理办法另行制定。 第四条 缴费单位应当到办理社会保险登记的社会保险经办机构办理缴费申报。 第五条 缴费单位应当在每月5日前,向社会保险经办机构办理缴费申报,报送社会保险费申报表(以下简称申报表)、代扣代缴明细表以及社会保险经办机构规定的其他资料。 缴费单位到社会保险经办机构办理社会保险缴费申报有困难的,经社会保险经办机构批准,可以邮寄申报。邮寄申报以寄出地的邮戳日期为实际申报日期。 第六条 缴费单位因不可抗力因素,不能按期办理社会保险缴费申报的,可以延期办理。但应当在不可抗力情形消除后立即向社会保险经办机构报告。社会保险经办机构应当查明事实,予以核准。 第七条 社会保险经办机构应当对缴费单位送达的申报表和有关资料进行即时审核。对缴费单位申报资料齐全、缴费基数和费率符合规定、填报数量关系一致的申报表签章核准;对不符合规定的申报表提出审核意见,退缴费单位修正后再次审核;对不能即时审核的,社会保险经办机构应当自收到缴费单位申报表和有关资料之日起,在最长不超过2日内审核完毕。 第八条 缴费单位不按规定申报应缴纳的社会保险费数额的,社会保险经办机构可暂按该单位上月缴费数额的百分之一百一十确定应缴数额;没有上月缴费数额的,社会保险经办机构可暂按该单位的经营状况、职工人数等有关情况确定应缴数额。缴费单位补办申报手续并按核定数额缴纳社会保险费后,由社会保险经办机构按照规定结算。 第九条 缴费单位必须在社会保险经办机构核准其缴费申报后的3日内缴纳社会保险费。缴费单位和缴费个人应当以货币形式全额缴纳社会保险费。 第十条 缴费单位必须按照条例第十二条的规定严格履行代扣代缴义务。缴费单位依法履行代扣代缴义务时,任何单位或个人不得干预或拒绝。 第十一条 缴费单位的缴费申报经核准后,可以采取下列方式之一缴纳社会保险费: (一) 缴费单位到其开户银行缴纳; (二) 缴费单位到社会保险经办机构以支票或现金形式缴纳; (三) 缴费单位与社会保险经办机构约定的其他方式。 履行前款规定的申报核准程序后,银行可以根据社会保险经办机构开出的托收凭证从缴费单位基本账户中划缴社会保险费。 第十二条 征收的社会保险费,应当进入社会保险经办机构在国有商业银行开设的社会保险基金收入户。社会保险经办机构应当按照有关规定定期将收到的基金存入财政部门在国有商业银行开设的社会保障基金财政专户。 第十三条 社会保险经办机构对已征收的社会保险费,根据缴费单位的实际缴纳额(包括代扣代缴额)、代扣代缴明细表和有关规定,按以下程序进行记账: (一) 个人缴纳的基本养老保险费、失业保险费和基本医疗保险费,分别计入基本养老保险基金、失业保险基金和基本医疗保险基金,并按规定记录基本养老保险和基本医疗保险个人账户;

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三、项目简介 发展新能源汽车是国际共识和我国的国家战略,电动汽车是主要技术选择。动力电池系统是电动汽车的技术瓶颈,其精准管理是保障整车高效、安全和动力电池长寿命运行的核心,动力电池状态量的高精度、强鲁棒性估计一直是行业技术攻关的国际难题和学术研究的前沿热点。项目组在国家自然科学基金、863计划等支持下,历时9年理论研究,取得系统性、原创性成果。 1.发现了动力电池输出特性与其内部参数和状态间具有的间接映射规律,阐释了端电压的动、静态分量解耦机制以及动态分量具有的多阶RC解析特性,建立了具有普适性的动力电池N阶电气解析模型,融合电化学机理模型、分数阶阻抗模型,建立了动力电池机理-频域-电气特性综合解析模型。发明了基于存档和实时运行数据驱动的五步骤动力电池模型参数辨识方法,实现了模型结构和阶次优化,大幅提高了模型的高精度、环境感知和可靠性等属性。 2.发现了动力电池荷电状态SOC与其静态电势/开路电压OCV相关且存在单调映射关系,建立了融合动力电池N阶电气解析模型的OCV在线辨识和SOC估计方法。针对OCV-SOC映射关系的差异性,首次提出了基于实车片段数据的SOC 映射参数重构方法。提出了将动力电池SOC和动态极化电压作为状态量、将输出电压作为观测量的SOC估计模型,增强闭环校正精度,建立了滤波器类动力电池自适应SOC估计算法体系。发明了多时间尺度动力电池SOC、容量、内阻的多状态量协同估计和精度自适应标定方法,提高了状态估计的鲁棒性。 3.发现了动力电池组系统充放电末期具有的强极化非线性特性并引发端电压明显的不一致性,首次提出分别以能量最大利用、安全应用为目标约束的动力电池组系统SOC分段高精度估计准则。揭示了动力电池组系统实际工作环境的差异对其性能衰退的影响机制以及动力电池不一致性对其性能衰退的耦合效应和演化机理,建立了以SOC、容量、温度等为自变量、以偏差为因变量的近似模型。首次提出基于“表征单体模型+偏差量化模型”的动力电池组系统状态估计算法,提高了状态估计的精度和适用性。 该项目在动力电池管理核心模型和算法方面做出原创性国际引领贡献,发表论文SCI检索49篇、ESI高被引21篇。8篇代表作SCI他引1130次、谷歌学术他引1956次、最高单篇他引504次,其中3篇入选“中国百篇最具影响学术论文”(电动汽车领域仅有的3篇)、2篇入选SCI期刊年度最佳论文奖。授权发明专利6件。完成人作国际学术会议特邀报告16次,任IEEE电气化交通大会ITEC2014和国际应用能源大会ICAE2016大会主席,入选科睿唯安高被引科学家。成果得到Shixue Dou、欧阳明高、Stefan Pischinger院士等国际学术同行的高度评价。项目成果在华为、北汽新能源、郑州宇通客车、联合汽车电子等企业得到成功应用,主要指标同比优势明显。中国汽车工程学会组织科技成果评价,认为“……建立了动力电池系统集成参数辨识与状态在线估计综合优化方法,动力电池状态估计技术处于国际领先水平”。

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